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The extinct order Coloneurodea [Insecta: Pterygota: Panorthoptera]: wing venation, systematics and phylogenetic relationships

100%

Bethoux O., Nel A., Lapeyrie J.

Annales Zoologici

2004

tom 54

nr 2

A new interpretation of the wing venation of the fossil insect order Caloneurodea is proposed. Complex fusions of veins are demonstrated, supporting the assignment of the Caloneurodea to the Panorthoptera, i.e. the order is closely related to the Orthoptera. A new diagnosis of the order is proposed. The following taxa are re-described after new interpretation, material or preparation, or are newly described: Caloneura dawsoni Brongniart, 1885 (= Boltonaloneura subtilis (Bolton, 1925) syn. nov.), Gigagramma carpenteri gen. and sp. nov.; Sthenarocera pachytyloides Brongniart, 1885, Pruvostiella lecomtei (Pruvost, 1919), Apsidoneura flexa Carpenter, 1943, Homaloptila similis (Meunier, 1911), Pleisiogramma medialis Carpenter, 1943 (=Pleisiogramma reducta Carpenter, 1943 syn. nov.), Paleuthygramma acuta Carpenter, 1943, Paleuthygramma sharovi sp. nov., Euthygramma parallelum Martynov, 1928 (= Euthygramma curvatum Martynov, 1931 syn. nov.; = Paleuthygrammella aberrans (Martynov, 1938) syn. nov.), Anomalogramma parva Carpenter, 1943, Nanogramma gandi gen. and sp. nov., and Ligogramma sinuosa gen. and sp. nov. The family Gelasopteridae Carpenter, 1976 is revised and its ordinal relationships are discussed. After the proposed phylogenetic frame, mutation(s) occurring in homeotic genes are expected to be responsible of the unusual wing morphology of the Caloneurodea, mixing fore- and hind-wing 'panorthopteriď features.

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Revision of the enigmatic insect family Anthracoptilidae enlightens the evolution of Palaeozoic stem-dictyopterans

88%

Guan Z., Prokop J., Roques P., Lapeyrie J., Nel A.

Acta Palaeontologica Polonica

2016

tom 61

nr 1

The position of the Palaeozoic Anthracoptilidae has been a major problem of insect systematics for over a century. The previous hypotheses suggested affinities of Anthracoptilidae with the Palaeodictyopteroida, Protorthoptera, Hypoperlida, stem-Mantodea, Paraneoptera, Holometabola, or Eoblattida. Herein we put forward a new hypothesis based mainly on a comprehensive re-evaluation of the wing venation characters and re-examination of the type material of Anthracoptilus perrieri and Mesoptilus dolloi. The Anthracoptilidae are considered as belonging to the paoliid lineage, sister group of the Dictyoptera. In particular this result refutes the presence of Mantodea in the Paleozoic fossil record. The two families Strephocladidae and Strephoneuridae, are considered as junior synonyms of the Anthracoptilidae, while the previous synonymy of the Anthracoptilidae with the Ischnoneuridae is rejected. We consider the Permarrhaphidae, previously considered as synonym of the Anthracoptilidae, (and Permarrhaphus) as Insecta incertae sedis. The following new taxa are proposed: Mesoptilus carpenteri sp. nov. from the early Permian of Wellington Formation in Oklahoma (USA) which extends the range and distribution of the genus; Pseudomesoptilus gen. nov. is designated to include Mesoptilus sellardsi; Strephocladus permianus sp. nov. from the middle Permian of South of France; Westphaloptilus gallicus gen. et sp. nov. from the Bashkirian of the North of France.

Ograniczanie wyników

1 Acta Palaeontologica Polonica

1 Annales Zoologici

2 Lapeyrie J.

2 Nel A.

1 Bethoux O.

1 Guan Z.

1 Prokop J.

1 Roques P.

1 2016

1 2004

Wyniki wyszukiwania

The extinct order Coloneurodea [Insecta: Pterygota: Panorthoptera]: wing venation, systematics and phylogenetic relationships

Revision of the enigmatic insect family Anthracoptilidae enlightens the evolution of Palaeozoic stem-dictyopterans