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The exquisitely preserved longipterygid enantiornithine Rapaxavis pani is redescribed here after more extensive preparation. A complete review of its morphology is presented based on information gathered before and after preparation. Among other features, Rapaxavis pani is characterized by having an elongate rostrum (close to 60% of the skull length), rostrally restricted dentition, and schizorhinal external nares. Yet, the most puzzling feature of this bird is the presence of a pair of pectoral bones (here termed paracoracoidal ossifications) that, with the exception of the enantiornithine Concornis lacustris, are unknown within Aves. Particularly notable is the presence of a distal tarsal cap, formed by the fusion of distal tarsal elements, a feature that is controversial in non−ornithuromorph birds. The holotype and only known specimen of Rapaxavis pani thus reveals important information for better understanding the anatomy and phylogenetic relationships of longipterygids, in particular, as well as basal birds as a whole.
Exceptionally well−preserved embryonic skulls of Upper Cretaceous (Campanian Anacleto Formation) sauropods from Auca Mahuevo (Neuquén Province, Argentina) provide important insights into the ontogeny and evolution of titanosaurian neosauropods. The most important cranial modifications occurring during titanosaurian ontogeny appear to be centered on the infraorbital and narial regions, which exhibit a substantial degree of “mosaic” evolution. On one hand, the Auca Mahuevo embryos show a large jugal that forms part of the lower margin of the skull and unretracted external nares, as indicated by the position and orientation of the lacrimals as well as the anterior extension of the frontals. Both of these features are ancestral for neosauropods, being present in prosauropods. On the other hand, the embryonic skull exhibits a large ventral notch, tentatively interpreted as homologous to the neosauropod preantorbital fenestra, that opens ventral to the jugal and between the maxilla and the quadratojugal, and a temporal region that closely resembles the adult neosauropod condition. This mosaic of character states indicates that different regions of the skull of titanosaurian neosauropods acquired their characteristic morphology at substantially different rates during their ontogenetic development.
Although recent discoveries from Lower Cretaceous sediments in northeastern China have greatly improved our understanding of the initial stages of avian diversification in eastern Asia, the early evolution of Aves elsewhere on the continent remains poorly understood. In 2004, a collaborative field effort directed by personnel from the Chinese Academy of Geological Sciences and Carnegie Museum of Natural History recovered multiple partial to nearly complete avian skeletons from outcrops of the Lower Cretaceous Xiagou Formation exposed in the Changma Basin of northwestern Gansu Province, China. Here we describe a thrush−sized partial skeleton comprised of a fragmentary pelvic girdle and largely complete hind limbs. A phylogenetic analysis of 20 avian ingroup taxa and 169 anatomical characters places the specimen in Enantiornithes, and within that clade, in Euenantiornithes. When coupled with additional recent discoveries from the Changma Basin, the new skeleton improves our understanding of early avian evolution and diversification in central Asia.
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