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Populations of two species of woodland rodents were studied: Apodemus flavicollis (A. f.) and Clethrionomys glareolus (C. g.)t inhabiting a set of small wood patches, isolated from large, continuous forest. The species composition, density and population dynamics differed from those in the forest. The rodents used the entire area as a patchy habitat, moving between the woodlots. In the breeding season high mobility caused higher mortality among males, especially in C. g. Sex ratio in C. g. was female biased. In A. f. females prevailed in spring whereas there was a prevalence of males in autumn. Seasonal changes in age structure followed different patterns in females and males. Males prevailed in first spring litters in both species. Males also prevailed among numerous immigrants of A. f. but females prevailed among immigrants of C. g. The demographic processes in these species resulted from habitat fragmentation and different life strategies.
Live-trapping of rodents was conducted over a 12-ha plot on unevenly spaced trap lines, with inner lines forming a 3.5-ha grid of closely spaced traps. This design was used to estimate the probability that bank voles Clethrionomys glareolus (Schreber, 1780) and yellow-necked mice Apodemus flavicollis (Melchior, 1834) trapped in small grids are true residents rather than visitors from the surrounding area. On average, 12% of the voles and 19% of the mice were trapped within and beyond the grid in the same trapping sessions. As these were mainly wide-ranging males moving over the plot, home ranges of males may be underestimated on small grids. In total, 36% of voles and 39% of mice that were marked in spring-summer were trapped at least once in their life beyond the grid. Typically, these were individuals shifting home ranges and migrating to or from the grid. The size of lifetime ranges of rodents was significantly larger than temporary home ranges and may therefore be underestimated on small grids. "Single­-capture" individuals were mainly true transients rather than visitors. Only 12% of voles and 15% of mice resided on the plot for longer time than in the grid.
Spatial behaviour of female voles Clethrionomys glareolus (Schreber, 1780) was studied for 7 years in woodland of Kampinos National Park, Poland. Annual peaks ranged from 25 to 40 voles per ha. Twenty-six percent of newly marked females were already fully-grown adults. Immature females were slightly less site-tenacious than mature ones. The total movement range does not differ much between immature and mature recruits. Home range shifts between successive trapping sessions were the longest in maturing females but also prevalent in pregnant or lactating ones. Non- maturing females as well as mature but not breeding ones were the least mohile and the most site-tenacious. The home range length and between-capture distances were greater for pregnant than for immature females. They were also greater for transients than residents. Extensive movements of reproducing females suggests that breeding dispersal may thus be much more widespread amongst Clethrionomys females than previously thought.
The reliability of the estimation of residency time of the bank vole Clethrionomys glareolus (Schreber, 1780) and the yellow-necked mouse Apodemus flavicollis (Melchior, 1834) in open populations was assessed by halving the 6-week intervals between trapping sessions. The status of "single-capture" individuals was identified to estimate the probability that they are native young rather than adult invaders. We found that the majority of "single-session" individuals constituted true transients that were present in the study plot for a short time only. They were young, immature rodents with a small body mass and a small number of captures. The probability that the "single-session" individuals stay on the plot for a longer time was 0.35 and 0.37 for voles and mice, respectively. These ostensible transients were trap-prone, fully-grown, mature adults, that revealed their presence on the plot already at the beginning of each trapping session. We found that "single-capture" individuals were mainly young, immature rodents that were not retrapped. The probability that the single-capture rodents are mature individuals, with a longer residency time, was 0.10 and 0.18 for voles and mice, respectively.
Population dynamics and spatial behaviour of the vole Clethrionomys glareolus (Schreber, 1780) and the yellow-necked mouse Apodemus flavicollis (Melchior, 1834) were studied for 7 years in woodland of Kampinos National Park, Poland. Mice were more mobile and less site-tenacious than voles. Annual peaks ranged from 9 to 104 voles and from 4 to 62 mice per ha. The highest densities of both species were preceded by winter breeding. The greatest movement range and the longest distance moved were observed in the years of low density, whereas in the high density year both species were least mobile and most site-tenacious. Intensive movements in the low density years led to early maturation, high turnover rates, and probably increased mortality. Low mobility and high site tenacity in the high density year enhanced population growth and suppressed maturation. Increasing density and cessation of breeding accounted for declining juvenile recruitment. Differences in movement pat­terns between years of low and high density were coupled with differences in the autumn age structure and winter mortality of both species.
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