Preferencje help
Widoczny [Schowaj] Abstrakt
Liczba wyników

Znaleziono wyników: 13

Liczba wyników na stronie
Pierwsza strona wyników Pięć stron wyników wstecz Poprzednia strona wyników Strona / 1 Następna strona wyników Pięć stron wyników wprzód Ostatnia strona wyników

Wyniki wyszukiwania

help Sortuj według:

help Ogranicz wyniki do:
Pierwsza strona wyników Pięć stron wyników wstecz Poprzednia strona wyników Strona / 1 Następna strona wyników Pięć stron wyników wprzód Ostatnia strona wyników
2
Artykuł dostępny w postaci pełnego tekstu - kliknij by otworzyć plik
Content available

Counting premolars in early eutherian mammals

100%
The primitive number of premolars for most eutherian groups is four. A growing number of Cretaceous taxa, however, had five. Regardless of the hypothesis used to explain the discrepancy, or what the primitive condition was, it is generally agreed that the middle (third) tooth of five-premolared taxa is the one not represented in mammals that have only four premolars. Hence the current practice of labeling the teeth as the first through fifth and the first through fourth, depending on how many teeth are observed in the jaw, results in incorrect implied homologies for the last two premolars of the series. Given the long-standing tradition of referring to the premolars as the first through fourth, for most eutherian groups, together with the uncertainties involved in interpreting the difference, the most practical solution is to refer to the disputed tooth by a neutral term, 'Px', as advocated several decades ago.
The Trinity therians have long been the focus of attempts to reconstruct the evolutionary history of higher mammals, especially in the context of the development of tribospheny. In this paper, we update the taxonomy of the tribosphenidan taxa known from the Trinity Group and establish with more confidence the premolar/molar count in each. Many isolated specimens can be referred to a specific tooth locus. Additional diversity is revealed within the Deltatheroida, with the description of an additional species of Oklatheridium; Pappotherium is here considered a likely metatherian based on the inferred presence of four molars, while Holoclemensia is a basal eutherian (the opposite of some traditional interpretations). The remainder of the genera, Kermackia and Slaughteria, cannot be allied with either of the living groups of tribosphenidan mammals using the available data. We identify strong morphological diversity within this assemblage of stem taxa, including modifications to the traditional tribosphenic occlusal pattern in Kermackia. Mammalian evolution at the base of the tribosphenidan radiation was complex, and this underscores the need for caution when interpreting the morphology and relationships of taxa known by incomplete material.
Polyglyphanodon sternbergi Gilmore, 1940 is a large−bodied lizard from the Late Cretaceous of North America distinguished by its transversely oriented, interlocking teeth. Initially the teeth of P. sternbergi were described as smooth and blade−like, but recent discoveries of new specimens from the type locality and re−examination of the original material indicate that the chisel−like teeth of P. sternbergi have small, irregular serrations along the blades. These serrations are similar in size to those found on the teeth of the modern herbivorous lizard Iguana iguana and were likely used in a similar manner to crop vegetation, but was also capable of a degree of oral food processing due to the transverse orientation and interlocking arrangement of the dentition of P. sternbergi. Additionally, the presence of transversely oriented teeth with V−shaped blades in the anterior portion of the tooth row of P. sternbergi represents an additional shared characteristic in tooth structure between P. sternbergi and Dicothodon moorensis, Bicuspidon numerosus, and Peneteius aquilonoius; all transversely−tooth polyglyphandontine lizards from the Cretaceous of North America. It appears that the unique dentitions of Polyglyphanodon sternbergi (large teeth with transverse, serrated blades) and Peneteius aquilonius (small teeth with mammal−like specializations) present by the end of the Cretaceous were derived from a bicuspid, transversely oriented precursor tooth with a V−shaped blade.
We describe a new boreosphenidan mammal, Atokatheridium boreni gen. et sp. n., from the Early Cretaceous of Oklahoma, based on an upper molar and a tentatively referred lower molar. The upper molar is characterized by a small protocone and unwinged conules, broad stylar shelf, paracone taller than metacone, and lack of pre- and postcingula. Comparisons with relevant Early and Late Cretaceous boreosphenidans suggest closest similarity to Deltatheroida, including one character (extreme development of the distal stylar shelf, which projects labially and lacks cusps) interpreted as derived. The tentatively attributed lower molar shows similarity to Deltatheridium and the ?aegialodontid genus Kielantherium in having the paraconid higher than the metaconid, but differs from Kielantherium in having a differently shaped talonid. From Aegialodon it differs in having a vertically oriented (rather than semi-procumbent) paraconid and a larger talonid. We figure also two isolated trigonids, differing in size, which show some resemblance to that of ?Atokatheridium. Deltatheroidans, despite their generally primitive dental morphology, are otherwise surely known only from the Late Cretaceous, and are largely restricted to the Old World. If a deltatheroidan, the new taxon implies a significant temporal range extension for the group, and provides another biogeographic link between Cretaceous mammals of Asia and North America.
Herein we describe the oldest well-sampled multituberculate assemblage from the Cretaceous of North America. The fauna is dated at 98.37 Ma and thus approximates the Albian-Cenomanian (Early-Late Cretaceous) boundary. The multituberculate fauna is diverse. Two of the multituberculates (Janumys erebos gen. et sp. n. and an unidentified taxon) are provisionally placed among 'Plagiaulacida'. Another taxon, Ameribaatar zofiae gen. et sp. n., is of uncertain subordinal affinities. The remaining multituberculates appear to represent the advanced suborder Cimolodonta and fall within the 'Paracimexomys group'. We rediagnose Paracimexomys on the basis of the type species, P. priscus, and refer to other species as cf. Paracimexomys (including cf. P. perplexus sp. n.). A revised diagnosis is also provided for Cenomanian Dakotamys. A previously-described species from the Cedar Mountain Formation is placed in Cedaromys gen. n. as C. bestia, together with C. pawus sp. n. Bryceomys is represented in the fauna by B. intermedius sp. n. Relationships of Paracimexomys-group to later taxa remain obscure. However, Bryceomys and Cedaromys share a number of features with Cimolodontidae. Given these resemblances, together with the fact that Cimolodontidae retain certain plesiomorphies (stout lower incisor, gigantoprismatic enamel) with respect to Ptilodontoidea (to which they are commonly referred), we suggest that Cimolodontidae may have arisen from a clade within the 'Paracimexomys group', independent of ptilodontoids.
7
51%
Mammalian diversity in North America shifted significantly during the Early Cretaceous, from archaic groups dominant in the well-sampled faunas of the Late Jurassic to advanced forms (including early members of modern clades) by the Albian–Cenomanian. However, the dynamics of this transition are poorly understood, since faunas of earliest Cretaceous age are unknown. Here we describe the first fossil mammals from exposures of the Lakota Formation in the Black Hills of South Dakota, a unit correlated with the upper Berriasian–lower Barremian and positioned stratigraphically between the underlying Morrison Formation and Aptian–Albian units exposed elsewhere in North America. The mammalian fauna from the Lakota Formation is transitional with regard to the North American fossil record, representing a broad spectrum of both Jurassic and Cretaceous lineages: present are “plagiaulacidan” multituberculates allied with Late Jurassic Allodontoidea and Early Cretaceous Plagiaulacoidea; the geologically youngest dryolestoid(s) and “triconodontine” triconodontids (characteristic Late Jurassic taxa from the Morrison Formation); the oldest spalacotheriid “symmetrodont”; the first record of an amphitheriid-like stem zatherian from North America (abundant in the Middle Jurassic–earliest Cretaceous of Europe); and the oldest North American tribosphenic mammal (abundant and diverse on the continent by the end of the Early Cretaceous). Taxa making their first North American appearance in the Lakota Formation (Plagiaulacoidea, including a genus also known from the Purbeck of Britain; Spalacotheriidae, stem Zatheria, Tribosphenida) are also known from the Early Cretaceous of Western Europe, suggesting the possibility that they represent immigrants.
The lowest part of the Monarch Mill Formation in the Middlegate basin, west-central Nevada, has yielded a middle Miocene (Barstovian Land Mammal Age) vertebrate assemblage, the Eastgate local fauna. Paleobotanical evidence from nearby, nearly contemporaneous fossil leaf assemblages indicates that the Middle Miocene vegetation in the area was mixed coniferous and hardwood forest and chaparral-sclerophyllous shrubland, and suggests that the area had been uplifted to 2700–2800 m paleoaltitude before dropping later to near its present elevation of 1600 m. Thus, the local fauna provides a rare glimpse at a medium- to high-altitude vertebrate community in the intermountain western interior of North America. The local fauna includes the remains of fish, amphibians, reptiles, birds, and 25 families of mammals. Carnivorans, the focus of this study, include six taxa (three of which are new) belonging to four families. Canidae are represented by the borophagine Tomarctus brevirostris and the canine Leptocyon sp. indet. The earliest record and second North American occurrence of the simocyonine ailurid Actiocyon is represented by A. parverratis sp. nov. Two new mustelids, Brevimalictis chikasha gen. et sp. nov. and Negodiaetictis rugatrulleum gen. et sp. nov., may represent Galictinae but are of uncertain subfamilial and tribal affinity. The fourth family is represented by the felid Pseudaelurus sp. indet. Tomarctus brevirostris is limited biochronologically to the Barstovian land mammal age and thus is consistent with the age indicated by other members of the Eastgate local fauna as well as by indirect tephrochronological dates previously associated with the Monarch Mill Formation. Actiocyon parverratis sp. nov. extends the temporal range of the genus Actiocyon from late Clarendonian back to the Barstovian. The Eastgate local fauna improves our understanding of mammalian successions and evolution, during and subsequent to the Mid-Miocene Climatic Optimum (~14–17 Ma).
9
51%
The description of a new genus (Apeomyoides) of eomyid rodent from the Miocene of Nevada increases the diversity of known taxa, enhances the geographic range, and extends the biochronology for the Apeomyinae (Eomyidae). Three groups of Eomyidae are known from the fossil record of North America. Of the three groups, Neogene taxa include four genera representing the Eomyinae and two genera representing the Apeomyinae; no genera of the subfamily Yoderimyinae are known from the Neogene of the continent. This diversity represents a significant reduction of eomyid taxa compared to the Paleogene, from which 17 genera of eomyines and three genera of yoderimyines are known. In Eurasia, 11 genera of eomyids occurred during the Neogene, with a few taxa that persisted until about 2 million years before present. At present, there are no known eomyids from the last 4.5 million years of the Neogene in North America. Apeomyoides savagei is referable to the subfamily Apeomyinae based on several key structures of the teeth and mandible. This new eomyid is part of the Eastgate local fauna, collected from volcanic ash deposits of the Monarch Mill Formation, Churchill County, Nevada. Apeomyoides has an occlusal pattern that shares characteristics of apeomyines from both North America (Megapeomys and Arikareeomys) and Eurasia (Apeomys and Megapeomys). The unique occlusal pattern and large size of Apeomyoides demonstrates that not all eomyids from North America were small or that their lineages decreased in size through time. Apeomyoides also may provide evidence, which challenges the hypothesis that eomyids within a single lineage from North America became more lophodont in geologically younger genera.
Sauroposeidon proteles is a large brachiosaurid sauropod recently described from the Antlers Formation (Aptian-Albian) of southeastern Oklahoma. Sauroposeidon represents the culmination of brachiosaurid trends toward lengthening and lightening the neck, and its cervical vertebrae are characterized by extensive pneumatic structures. The elaboration of vertebral air sacs during sauropod evolution produced a variety of internal structure types. We propose a new classification system for this array of vertebral characters, using computed tomography (CT) of pneumatic internal structures. Comparisons with birds suggest that the vertebrae of sauropods were pneumatized by a complex system of air sacs in the thorax and abdomen. The presence of a thoraco-abdominal air sac system in sauropods would dramatically affect current estimates of mass, food intake, and respiratory requirements. Sauroposeidon was one of the last sauropods in the Early Cretaceous of North America; sauropods disappeared from the continent by the early Cenomanian. The demise of sauropods in the Early Cretaceous of North America predates significant radiations of angiosperms, so the decline and extinction of this dinosaur group cannot be linked to changes in flora.
11
51%
The Middle Jurassic mammal Shuotherium has lower molars that possess a trigonid and talonid, but are unique in having the talonid situated in front of the trigonid, rather than behind it, as in molars of usual tribosphenic pattern. Shuotherium dongi Chow and Rich, 1982 was based on a dentary bearing seven teeth, originally interpreted as three premolars and four molars. Based on comparison with other groups of early mammals, we reinterpret the premolar–molar boundary in the holotype of S. dongi, and propose a dental formula of four (or more) premolars and three molars. The ultimate lower premolar (previously identified as the first molar) has a completely developed trigonid and no talonid or pseudo−talonid. We hypothesize that the mesial cingulid on molars of Australosphenida is a highly plausible structural antecedent to the pseudo−talonid of Shuotherium. This and other shared, derived features support a relationship of Shuotherium and Australosphenida as sister−taxa. We hypothesize that the common ancestor of Shuotherium + Australosphenida had a global distribution no younger than early Middle Jurassic, and that the respective clades diverged prior to full separation of Gondwanan and Laurasian landmasses.
12
Artykuł dostępny w postaci pełnego tekstu - kliknij by otworzyć plik
Content available

In quest for a phylogeny of Mesozoic mammals

51%
We propose a phylogeny of all major groups of Mesozoic mammals based on phylogenetic analyses of 46 taxa and 275 osteological and dental characters, using parsimony methods (Swofford 2000). Mammalia sensu lato (Mammaliaformes of some authors) are monophyletic. Within mammals, Sinoconodon is the most primitive taxon. Sinoconodon, morganucodontids, docodonts, and Hadrocodium lie outside the mammalian crown group (crown therians + Monotremata) and are, successively, more closely related to the crown group. Within the mammalian crown group, we recognize a fundamental division into australosphenidan (Gondwana) and boreosphenidan (Laurasia) clades, possibly with vicariant geographic distributions during the Jurassic and Early Cretaceous. We provide additional derived characters supporting these two ancient clades, and we present two evolutionary hypotheses as to how the molars of early monotremes could have evolved. We consider two alternative placements of allotherians (haramiyids + multituberculates). The first, supported by strict consensus of most parsimonious trees, suggests that multituberculates (but not other alllotherians) are closely related to a clade including spalacotheriids + crown therians (Trechnotheria as redefined herein). Alternatively, allotherians can be placed outside the mammalian crown group by a constrained search that reflects the traditional emphasis on the uniqueness of the multituberculate dentition. Given our dataset, these alternative topologies differ in tree−length by only ~0.6% of the total tree length; statistical tests show that these positions do not differ significantly from one another. Similarly, there exist two alternative positions of eutriconodonts among Mesozoic mammals, contingent on the placement of other major mammalian clades. Of these, we tentatively favor recognition of a monophyletic Eutriconodonta, nested within the mammalian crown group. We suggest that the “obtuse−angle symmetrodonts” are paraphyletic, and that they lack reliable and unambiguous synapomorphies.
Pierwsza strona wyników Pięć stron wyników wstecz Poprzednia strona wyników Strona / 1 Następna strona wyników Pięć stron wyników wprzód Ostatnia strona wyników
JavaScript jest wyłączony w Twojej przeglądarce internetowej. Włącz go, a następnie odśwież stronę, aby móc w pełni z niej korzystać.