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Iberovaranus Hoffstetter, 1969 was erected as a monotypic genus of varanine varanid lizard on the basis of a single trunk vertebra from the Miocene of Spain. Thanks to the study of the holotype, as well as of a still undescribed cervical vertebra from the same locality, we show that the vertebral morphology of Iberovaranus is contained within the known variability of Varanus. Therefore, Iberovaranus Hoffstetter, 1969 is considered a subjective junior synonym of Varanus Merrem, 1820, and the species Iberovaranus catalaunicus Hoffstetter, 1969 should be considered a nomen dubium.
A cladistic analysis of Megadontosuchus arduini from the middle Eocene of Monte Duello (NE Italy) confirms its tomistomine relationships, but the low number of scorable characters determines a low resolution within the tomistomine clade. However, Megadontosuchus is clearly distinct from the other Eocene European or North African tomistomines, in having a moderate elongated but robust rostrum, massive maxillary and dentary teeth and large supratemporal fenestrae. The rostrum and teeth characteristics could indicate that M. arduini had a degree of feeding specialization intermediate between Maroccosuchus zennaroi and the Eocene European tomistomines. A summary of tomistomine palaeobiogeography suggests that despite only one species with a rather restricted range survives at present, such a clade had a glorious past with a world wide distribution documented by a conspicuous fossil record that starts at least in the early Eocene. At present, a detailed knowledge of tomistomine palaeobiogeography is hindered by the lack of modern taxonomic revisions of some fossil remains and therefore by the poor understanding of phylogenetic relationships.
Originally designated by Dollo in 1907, the holotype of Eosuchus lerichei has never been carefully described but simply cited and compared in a number of papers. This work is an attempt to fill this gap and to place this taxon in a cladistic phylogenetic context. E. lerichei can be considered a valid basal gavialoid from late Paleocene of North Western Europe, sharing the presence of extremely enlarged foramina aerea on quadrates with the coeval Eosuchus minor from eastern North America (formerly described as Gavialis minor). These two species can be considered sister taxa and, for priority reason, they should be both ascribed to genus Eosuchus. The results of the cladistic analysis show that the European species possess characters that can be considered as slightly derived if compared to those of its American relative, suggesting an eastward dispersion from North America before the Paleocene–Eocene boundary and before the full opening of the Atlantic Ocean or local evolution from a basal gavialoid stock similar to E. minor. Both species of Eosuchus come from marine outcrops and represent a further evidence for the salt−water tolerance of the earliest stages of Gavialoidea evolutionary history. Despite the present endemicity of the only living gharial, Gavialis gangeticus, the historical biogeography of gavialoids shows a lost global distribution and reveals several transoceanic dispersals.
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