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The hyolith assemblage from the Lower Cambrian Bastion Formation of North−East Greenland is significant in that it contains several hyolith taxa that possess traits of both orders Hyolithida and Orthothecida. They possess morphological traits that seem to be characteristic of the ancestral forms of both groups. In addition, many hyolith taxa from this interval are globally distributed, supporting the notion that these fossils have potential as stratigraphic indicators. This assemblage contains genera and/or species seen in Australia, North America, the Siberian Platform, and South China. Hyoliths identified include the hyolithids Parkula bounites, Hyptiotheca karraculum, Microcornus eximius, M. petilus, Paracornus poulseni gen. et sp. nov., as well as Similotheca similis?, S. bastionensis sp. nov., and S. groenlandica sp. nov.; two opercula remain in open nomenclature. Orthothecids from this assemblage are one unnamed species each of Contitheca and Gracilitheca. Large, macro−sized hyoliths from the same formation described by Poulsen (1932) are mostly unidentifiable, although an operculum formerly identified as Hyolithes (Orthotheca) communis is reassigned to Hyptiotheca. Problematic organisms of uncertain affinity include Cupitheca holocyclata, Conotheca australiensis, an unnamed species of Coleolus, and the cap−shaped Cassitella baculata gen. et sp. nov. that may be an operculum of some as yet unknown organism. Missarzhevsky (1969) used Hyolithes(Orthotheca) bayonet var. groelandicus and H. (O.) bayonet var. longus as the basis for Lenatheca, but the specimens on which that genus is based are too poorly known for a proper diagnosis of Lenatheca.
Newly discovered specimens of the orthothecide hyoliths in Hebei Province of North China reveal new data on diversity, skeletal microstructure and palaeoecology of the order Orthothecida. Decoritheca cyrene possesses a well-defined skeletal structure consisting of rows of tubercles in the shell wall, which correspond to dimples on the internal surface of the conch. We describe a new species Cupitheca convexa sp. nov. characterised by a planar and interior convex operculum with a pair of bilobate cardinal processes, which differs from the more widespread C. holocyclata in opercular morphology and overall ornamentation of the conch, highting the significance of operculum in the classification of hyoliths. First discovery of the presumed unusual protoconch of C. convexa sp. nov. implies a possible lecithotrophic development to adapt to nutrient-fluctuant environments. Pitting structure on the operculum and shell as well as lamellar- fibrillar structure on the internal mould of C. convexa sp. nov. are observed herein.
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