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Actual status of mountain forests especialy, of Norway spruce (Picea abies Karst. (L)) stands is a result of combined action between forest management measures and biotic and abiotic risk factors. In this context, tree growth dynamics is a result of multifactor interaction (competition, disturbance, management practice, climate etc.). Wind damage affect the stand structure and normal ecosystem functions including the relationship between individual tree growth and competition processes. Though, it is recorded a diminishing of productive capacity for affected stands as well as a significant diameter increment for remaining standing trees. The correlation between tree growth (cumulate basal area increment in the last 10 years) and competition (Hegyi and Schutz competition indices) was analysed in stands with different windfall intensity. No relationship between tree growth and competition was observed in highly affected stands. Schutz competition index explains better individual tree growth comparing with Hegyi index in low or no-affected stand by windthrow. The variability of the basal area increment (lnΔɡ10), explained by competition indices, is rather low (<2%) in the stand that had been highly affected by windfall, regardless of the selected competition index. Thus, significant reduction of the stand density related to number of trees (low density), as a consequence of high intensity windfalls, crown competition has a very small influence on the basal area increment comparing with situations met in low (high density) and moderate (medium density) affected stands.
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Determination of the biomass of Pinus mugo stands

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The dwarf pine stands on unoriginal sites in mountainous areas of the Czech Republic are a current topic of scientific discussion. One of these sites is on the summits of the Hrubý Jeseník Mts. Various proposals for dwarf pine removal have been hindered by the absence of charts or tables that could be used to calculate how much biomass would need to be removed. Therefore, we created a methodology for dwarf pine biomass determination and applied it to five research transects of different ages. Based on the biomass estimates, we created trend curves illustrating the increase in biomass (dependent on age) as well as equations that could be used to roughly estimate the biomass of any dwarf pine stand, regardless of age or canopy level, for sites above the timberline in Hrubý Jeseník Mts. The equations for biomass calculations could also be applied to other mountain ranges where artificially planted dwarf pines of the same seed origin or the same morphological appearance as those existing in the Hrubý Jeseník Mts. are found.
A subarctic-boreal relict species, Salix lapponum, found within the area of the Łęczna-Włodawa Lakeland (Eastern Poland), is particularly threatened with extinction due to the southern limit of its range and the specific conditions of its occurrence. This conclusion is evidenced by a systematic reduction in the number of its locations, which is accompanied by a decrease in the numbers of individuals making up its populations. The aim of the present study was to determine habitat conditions of the occurrence of Salix lapponum populations on the basis of an analysis of physico-chemical factors of groundwater at the sites associated with various stages of succession taking place in small peatland water bodies. The results of this study confirm the trend that the stands and population numbers of this species are decreasing with the intensification of ecological succession and the degree of its progress and at the same time show that this species exhibits a wide amplitude of many of the investigated factors. Among the groundwater parameters under investigation, the low level of total nitrogen, phosphorus fraction and DOC as well as the high level of Ca, pH and electrolytic conductivity can be considered to be a set of conditions that promote the maintenance of Salix lapponum populations. On the other hand, in the case of nitrates, nitrites, sulphates as well as the Na, K and Mg ions, the Kruskal-Wallis statistical test showed that the distribution of their values was invariant in relation to the studied habitats, which is probably evidence that they do not determine the development or extinction of the investigated populations in these habitats.
Within the framework of work on the anatomical structure and durability features of wood, as well as analyses of radial growth, a preliminary analysis was carried out in regard to variation in the heartwood within Black locust trees. The research was carried out in three stands of straight-stemmed trees in western Poland. The variation displayed by the heartwood around its circumference and the eccentricity of the trunks was characterised with reference to: the heartwood radial index (HRI), the cross-sectional shape factor for heartwood (CSsf) and the pith eccentricity index (PEcc). The results confirmed a relationship between the degree of variation around the circumference of the heartwood and the mean age of the stands, while at the same time making clear the high level of differentiation in radial variation at the level of the individual tree. The findings suggest that the fertility of the habitat does not exert much of an influence on the generation of heartwood in Black locust trees. No statistically significant relationship was obtained when the circumferential variation of the heartwood was set against the total height or diameter at breast-height. Only in the case of crown length, and then only at the Wołów site, was there a moderate negative correlation with the coefficients of circumferential variation.
The studies carried out in the years 2004-2005 and in 2009 were aimed at identifying the functioning conditions of a boreal plant relict, Salix lapponum, in its stands under anthropopressure in the Łęczna-Włodawa Lakeland. The study results confirmed that the number of S. lapponum stands on Lake Miejskie and on Cow Marsh and its population size had significantly decreased as well as the population condition had deteriorated during the last 50 years. Based on the results of an analysis of environmental (biocenotic and abiotic) factors it was found that changes in habitat conditions in its stands were the main threat for downy willow. Another cause of population decline and the deteriorated condition of downy willow individuals may be the habitat isolation. The observed intraspecific phenomena, i.e. the lack of flowering in individuals and the lack of downy willow seedlings could be the reason for insufficient reproduction efficiency, ipso facto, the weaker adaptation of individuals to changes in habitat condition. Therefore, it seems that it is necessary to pursue active species conservation and protection which should primarily involve population reconstruction – restitution.
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Quality of Japanese larch stands in Poland

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The quality of 39 Japanese larch stands, mainly in north-western Poland, was characterized. On the basis of the presented results it can be concluded that properly managed forest stands of Japanese larch can compete in trees quality with stands of native larches. The alien species is more susceptible to a lower stand density and exposure to strong winds. This is no correlation between forest stand quality and growth dynamics.
Gap regeneration in a European beech (Fagus sylvatica L.) forest reserve was analyzed in relation to within-gap resource heterogeneity and ground vegetation competition. The study was carried out in two one-hectare permanent research plots (PRP) which included five smaller research plots (RP) encompassing two large gaps (500–700 m2), two small gaps (300–400 m2), and location under canopy. The coverage of woody regeneration, ground vegetation, dead wood, seedling density in eight height classes, characteristics of dominant trees of the beech regeneration, and the total thickness of holorganic horizons were measured. Soil moisture and light conditions were also assessed in selected sample plots. The relative direct and diffuse light was estimated by hemispherical photography. Small gaps showed both the highest cover of tree regeneration and the highest density of individuals per hectare. Slightly less regeneration was recorded in large gaps, while under closed canopy, regeneration densities were 5–10 times lower than in small gaps. Beech regeneration cover and the size (diameter and height) of dominant beech seedlings were positively related to relative diffuse light and negatively related to ground vegetation cover. The latter was positively related to diffuse light and soil moisture content. A pronounced statistically significant contrast in the cover and size of beech regeneration in relation to micro-site conditions (diffuse light, cover of graminoids) was only confirmed between sample plots located below canopy cover and those within gaps. Graminoids, in particular Calamagrostis epigejos L., occurred mainly in the large gap centre and along the southernmost edge of the large gap, increasing competition for resources here. The cumulative cover of ground vegetation and regeneration was relatively low (9–56%) compared with more mesic natural beech forests. The indicated negative influence of direct light at the northern gap edge suggests that extension of gaps on comparable sites in managed forest should not proceed in this direction.
The aim of the study was to determine the effect of pH and soil chemical composition on morphological traits of common tansy (Tanacetum vulgare L.). A two-year study was conducted based on observations, measurements and analyses of material collected from 24 stands in the following regions: Równina Włocławska (Włocławek Plain), Wyżyna Lubelska (Lublin Upland), Padół Zamojski (Zamość Depression), Działy Grabowieckie (Grabowiec Divide) Płaskowyż Suchedniowski (Suchedniów Plateau), Wysoczyzna Siedlecka (Siedlce High Plain) and Niecka Połaniecka (Połaniec Basin). Morphological traits of tansy shoots were determined: number of plants per clump, stem length, number of leaves per plant, total leaf length, petiole length, number of branches from the main stem, number of flower heads per corymb and fl ower head diameter. A soil analysis was performed determining the content of magnesium, potassium and phosphorus available for plants. The soil pH was also determined. The collected results indicate significant differentiation of the investigated samples, both in terms of morphological traits of the shoots and the chemical composition of the soils. A negative correlation was found between soil pH and number of leaves per shoot as well as number of branches per corymb. A positive correlation was noted between soil pH and total leaf length as well as between magnesium content and number of flower heads per corymb. The increased content of available phosphorus in the soil was associated with petiole shortening. A growth trend in the tansy shoot length was observed at larger amounts of available magnesium in the soil. On soils with low nutrient availability and a low pH, a reduced population density of the species in question was observed.
The paper presents the results of investigations into the variability of certain features of the wood macrostructure, such as the width of the annual ring, the share of late wood, and the relative density of the wood of grand fir, growing in four stands in southern Poland. The trees under investigation were 30-35 years old. The research material consisted of incremental cores sampled from trial trees with the use of a Pressler borer. The examined wood features were analysed in sections, each one comprising five annual rings. The rings were given numbers, sorted in an ascending order from the trunk circumference. The mean values of the analysed features were as follows: the annual ring width – 4.38 mm, the share of late wood - 35.39%, the relative wood density – 0.354g·cm-3. It was established that the wood of grand firs growing on less fertile soils had significantly narrower annual rings and higher wood density than those on more fertile soils.
It is not clear to what extent trees growing on anthropogenic sites change their growth and biomass allocation to different organs. We assessed the aboveground biomass in a chronosequence of six Scots pine monocultures (between 6 and 20 years old) to examine how precisely the site-specific or control allometric equations may reflect the value of biomass accumulation and allocation in stands growing in harsh site conditions on the overlayer spoil heap made as a result of opencast brown coal mining. The site is characterized by poor edaphic and water conditions and nutrient deficiency. The control equations were developed from Scots pine stands growing on post-agricultural lands in the close vicinity of the spoil heap. We found that equation type significantly influenced results of predicted biomass accumulation for all biomass components studied (although results were only marginally significant for total aboveground biomass, P=0.08). Total aboveground biomass in younger stands (6–9 years old) estimated using site-specific equations was >40% higher and for older stands (17–20 years old) from 7 to 27% lower than estimated using equations developed for the control sites. Our study revealed that under harsh environmental conditions in spoil heaps, biomass of young Scots pine stands significantly differ from values calculated based on control equations developed for more fertile soils with better water conditions in the same region. The control biomass equations may not be suitable to estimate biomass accumulation in stands growing on infertile habitats with poor water conditions, if the control equations are developed for nearby stands but growing under better site conditions.
The paper analyses the production and allocation of biomass in young, spontaneoussilver birch afforestation occurring on post-agricultural lands in the Mazowszeregion (central Poland). We investigated 114 sample plots of age varying from 1 to 19 years. During the first 15 years after their establishment on abandoned farmland, the naturally regenerated silver birch stands produced on average approximately 75 tons of dry biomass per hectare. The major (50–70%) part of this biomass was stored in the tree stems and this share increased with age. The fractions of biomass in the foliage and roots decreased over time, while the share of biomass in the branches remained rather constant. The significant age-dependency of the allometric relationships suggested the need to use age-sensitive biomass expansion factors to estimate the biomass from the stem volume.
Modern forestry management should be based on the principle of sustainability.In order to preserve habitat productivity, the amount of nutrients removed from the environment along with forestry products must be taken into consideration. This study shows the exact concentrations of chemical elements in different tree parts of Scots pine, growing on poor soils in north-western Poland. The observedvalues were compared to the values found in literature. In addition, the relationship between the concentrations of elements and stem diameter or stand density was researched. The highest concentration of chemical elements was observed in the needles (C, N, P, K, Mg, S, Mn, Na, Fe) and the lowest (C, N, P, S, Cu, Na, Ni, Pb, Zn, Fe) in the stem wood. Most of the macronutrients (P, K, Ca, Mg and S) reached optimal values, with the exception of N showing a deficiency, especially in the needles. The relationship between the content of elements and DBH or stand density was rather weak, and in both cases, negative.
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