The metabolites in the ovule come from the receptacle, ovary wall and placenta. They are accumulated in the chalaza and then translocated in reproductive and somatic structures via a system of specialized tissues. The hypostase is of great importance because it lies at the boundary of the chalaza with the nucellus and integuments, and contacts the vascular bundle. In the megagametophyte, the cell walls, which have many outgrowths, receive metabolites from the hypostase via podium and postament tissues. Histochemical data on accumulation of proteins, polysaccharides, lignin and tannins in tissues during ovule and seed development are presented. The dynamics of these substances and the character of metabolism in tissues are suggested as indicators of metabolite flow.
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