During the last two decades, an abundant selachian assemblage has been collected from the late Ypresian (NP12) fossiliferous sands of Prémontré (Aisne, northern France) but has received little attention. Sharks of the family Triakidae (Carcharhiniformes) are particularly well represented and all are described and figured herein. Among them, two new species of the genus Galeorhinus are described: G. duchaussoisi sp. nov. and G. louisi sp. nov.; these are compared to the common Paleogene G. ypresiensis which is refigured. Another triakid taxon, the genus Gomphogaleus gen. nov., is described. Most of the triakids have been recorded elsewhere in the North Atlantic region, suggesting a wider distribution than expected for these small sharks during the Paleogene. The present paper updates the list of selachians from Prémontré, bringing the number of taxa from 19 to 33 (including 22 sharks and 11 batoids) and improving our knowledge of the ancient North Atlantic Ypresian selachian fauna. Despite this vastly improved record, it is clear that fossil data are still very incomplete and insufficient for calibrating phylogenetic hypotheses of living forms. Review of the Prémontré fauna shows that the Triakidae were much more diverse and broadly distributed than at present, suggesting that the limited distribution and low diversity of living forms is probably a recent phenomenon.
Priabonian deposits from two localities, Kurgan and Derney, in the Trans−Urals (Western Siberia) have yielded numerous selachian teeth. The carcharhiniform family Scyliorhinidae is represented by three species, including two new: Premontreia uralica sp. nov., Foumtizia zhelezkoi sp. nov., and F. pattersoni (Cappetta, 1976). Both genera are recorded from the West−Siberian Basin for the first time. Presumably all three species (including the one morphologically mimicking F. pattersoni, a species recorded from Ypresian of England) are endemic for the Trans−Urals region (contrary to the cosmopolitan lamniforms recorded previously from the area. The known distribution pattern of extant scyliorhinids supports the probability of endemism of the cat sharks from the West−Siberian Basin, which has also been isolated geographically from the Peri−Tethys during the Late Eocene marine regression. The peculiar local environmental conditions due to the Priabonian climatic cooling in the Boreal realm might have also facilitated the speciation.
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