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During the Devonian the sedimentation on the continental shelves of Ardenne Massif and Boulonnais has changed from a mixed siliciclastic-carbonate ramp (Eifelian), through a carbonate barrier reef (Givetian) and then to a detritic influx with local mud-mounds (Frasnian). Here we analysed the faunistic dynamics of the trilobite associations through the changing environment. We used multivariate analyses (clustering and ordering) to discriminate the trilobite associations within 67 different samples. Three previously known communities and one new were recognised: the Eifelian Mixed association, the Givetian Dechenella association and the two Frasnian Bradocryphaeus and Scutellum-Goldius associations. These trilobite faunas present a progressive ecological specialisation. The Mixed association occurs both in the ramp or carbonated (local reef developed on the ramp) facies without any significant difference in its composition. The Dechenella fauna occurs preferentially close to barrier reefs, but can also survive during short periods of detrital input. The two Frasnian communities show a strong relationship with their environment. The Scutellum-Goldius association is only found in reef systems, whereas the Bradocryphaeus flourishes exclusively in lateral facies.
Previously, it was believed that there was a dramatic turn−over in the fauna of beavers (Rodentia, Castoridae) from the Barstovian to the Clarendonian from Nebraska. Stratigraphically controlled collections of fossil castorids from the Valentine Formation, which cross this boundary, now indicate that a complete change in the castorid fauna did not occur, but instead a more gradual change and replacement of earlier taxa with more advanced taxa occurred. The range of Eucastor tortus and Monosaulax skinneri is extended from late Barstovian into the latest Barstovian Devil’s Gulch Member and the range of the otherwise Clarendonian species of Prodipoides is extended downward into the late Barstovian Crookston Bridge Member. The skulls of Monosaulax skinneri and Eucastor valentinensis are described in detail for the first time. The cranial morphology of M. skinneri is primitive for Castoroidini and that of E. valentinensis is specialized for tooth−digging behavior as in the Nothodipoidini. A new genus, Temperocastor, is proposed for E. valentinensis based on its primitive morphology of the cheek teeth and derived cranial morphology. Temperocastor represents the most primitive nothodipoidine.
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