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In the Podolian Dniester Basin (southwestern Ukraine) the Lower Devonian marine deposits are represented by about 530 m thick continuous sequence of interlaminated carbonate and schale outcrops at several localities. Conodonts occur in most of the carbonate layers of the whole Lochkovian but are not abundant and their ramiform elements are mostly broken or lacking. Therefore, only the pectiniform, Pa elements of twenty five stratigraphically important conodont species occurring in the region are discussed and two new species, Caudicriodus schoenlaubi and Pandorinellina? parva are proposed. The hypothetical phyletic relationships within the main representatives of the icriodontid and spathognathodontid genera, Caudicriodus, Zieglerodina, and Pandorinellina? are traced. Comparison of the previously published and newly obtained data revealed discrepancies in the hitherto used interpretation of some of the conodont taxa and their stratigraphic ranges. Contrary to the earlier reports, Caudicriodus postwoschmidti does not occur in the lower Lochkovian but only in the middle part of the Chortkiv Formation, high above the Monograptus uniformis Zone. Based on new material and verification of the previous determinations, a modified scheme of the Lochkovian conodont zonation in Podolia is proposed. Conodont zones: Caudicriodus hesperius, C. transiens, C. postwoschmidti, C. serus, and ?Caudicriodus steinachensis are distinguished. The zones are correlated with conodont zonations in other regions—Barrandian, Cantabrian Mountains, Pyrenees, and Nevada. Biostratigraphy of the Siluro−Devonian transition and Lochkovian is integrated with the carbon isotope stratigraphy.
A cosmopolitan spathognathodontid taxon “Ancyrodelloides carlsi” is not regarded in this paper as one of the earliest known representatives of Ancyrodelloides according to former concepts, but is considered as conspicuous member of the genus Lanea. The taxon shows distinct morphological innovation of the upper surface of the Pa element within the Lanea lineage which makes it easily recognizable worldwide and suitable for global correlation. Presented stratigraphic correlation using conodonts and other faunal groups qualifies a short−lived Lanea carlsi as probably the best marker of the middle Lochkovian base. This paper presents reconstruction of the apparatus of L. carlsi which is the first complete reconstruction in the genus Lanea. The comparison of the Lanea apparatus and assumed apparatus of early Ancyrodelloides from the Požáry Quarries (Barrandian, Czech Republic) resulted to proposal of an alternative concept of evolution of these two genera. The relatively short−lived genus Ancyrodelloides is considered to split off from the Lanea clade in the late middle Lochkovian by the entry of Ancncyrodelloides transitans.
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Conodonts of the Upper Permian of Poland

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Eight species of conodonts, assigned to five genera, are described from the Zechstein limestone horizon of the Werra cyclothem from Wejherowo I.G.-1 boring in Pomerania. Lonchodina vistulensis n.sp., Prioniodina lindstroemi n.sp. and Hibbardella baltica n.sp. have been erected as new species. The Zechstein conodonts have been compared with the conodonts being parts of so-called natural assemblages.
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Conodonts, a large group of tiny extinct marine animals ranging in age from the Late Cambrian to Late Triassic (ca. 500 to 200 Mya), are usually considered as jawless vertebrates. Their only commonly occurring fossilized remains are minute, phosphatic, teeth−like elements of their feeding apparatuses. In most of the early conodonts the elements were conical and strongly elongated. Many of them are characterized by possession of a deep, longitudinal groove, usually associated with sharp edges or ridges. A comparative study of the grooved elements and venomous teeth and spines of living and extinct vertebrates strongly suggests that the groove in conodonts was also used for delivery of venom. Structural convergence of the conodont apparatus Panderodus with the grasping apparatus of chaetognaths, a group of extant, venomous invertebrate predators of similarly ancient origin, provides additional support for this conclusion.
Lenodus variabilis occurs in an interval in the lower to middle Kundan Stage which includes the Arenig–Llanvirn boundary. The lower part of its range constitutes the L. variabilis Zone,the upper part,where it co−occurs with Yangtze−placognathus crassus,making up the Y. crassus Zone. The conodont faunas of these two consecutive conodont biozones have been investigated in 14 sections from south,south−central and central Sweden,the faunal differences between the areas being mainly expressed in shifts in relative abundance. The taxon which is generally the most abundant, Baltoniodus, has comparable abundance maxima at localities representing shallow parts of the basin as well as in deeper parts,indicating that some environmental factor other than water depth has influenced its distribution. Drepanodus and Protopanderodus rectus prefer areas representing deeper parts of the epicontinental sea. Scalpellodus is most abundant at the sites representing intermediate depths. Semiacontiodus cornuformis was adapted to shallow water and is lacking at levels representing maximum depths, but at some places (for instance in Skåne) it is replaced by Semiacontiodus davidi.
A rich conodont fauna from two Neotethyan sections, Pizzo Mondello (western Sicily, Italy) and Pignola 2 sections (southern Apennines, Italy) includes conodonts described herein as a new species. Metapolygnathus praecommunisti sp. nov. is transitional between Paragondolella noah and Metapolygnathus communisti. The genus Metapolygnathus (including M. praecommunisti) is now characterised by posterior prolongation of the keel termination, associated with a centrally located pit and with a weak ornamentation confined to the anterior part of the platform margins. The establishment of M. praecommunisti addresses the problems related to the origin and the peculiar, probably facies−controlled, distribution of its descendant species M. communisti. Since M. praecommunisti occurs in the entire Tethys and in North America, we propose the species as a good guide fossil for global correlations, characterised by a short temporal range limited to the uppermost Tuvalian (upper Carnian). The stratigraphic occurrence of the genus Metapolygnathus is restricted to the Tuvalian–Lacian (upper Carnian–lower Norian), excluding its presence in the Julian substage (lower Carnian).
Well exposed Early–Middle Frasnian (E–MF, Palmatolepis transitans to Palmatolepis punctata zonal interval) deposits of the Holy Cross Mountains, in particular the reference Wietrznia section at Kielce, were studied in terms of conodont biofacies dynamics. Frequency of the conodont elements has been controlled mostly by depositional rate in hemipelagic muddy lithofacies and post−mortem gravity sorting during lateral redeposition in storm−generated, talus−like and encrinite layers. The conodont assemblages are dominated by a highly varying proportion of polygnathid, icriodontid, and ancyrodellid fauna. Major biofacies turnovers coincided with the deepening pulses corresponding to Timan, Middlesex, and early Rhinestreet global events. Trends in the conodont dynamics, mortality, and diversity point that the biotic shifts also coincide with the large−scale δ¹³C excursions as a record of changing trophic conditions during the major biogeochemical perturbation. A gradual decline of the Early Frasnian Ancyrodella reef−dwelling community correlates with the minor positive and succeeding larger negative δ¹³C excursion, and this is paired with a replacement by, mostly sparse, polygnathid and polygnathid−icriodontid biofacies, as well as with a short−term Belodella acme in mud−mounds areas. The distinctive habitat deterioration in pelagic and reef ecosystems is broadly correlative with the Domanik Crisis. The progressive biofacies unification is a conodont response to onset of the prolonged (ca. 0.5 Ma) δ¹³C enrichment, probably linked with high−stress life conditions due to eutrophication and partly anoxic regimes. A negative carbon isotope excursion in the late Pamatolepis punctata Zone is marked by the second major biofacies turning point during the Rhinestreet transgression, as recorded primarily in a final mesotaxid extinction, and highlighted also by decrease of conodont size and increased mortality of juveniles. After stabilization of δ¹³C values and a return to the background level across the Palmatolepis punctata–Palmatolepis hassi zonal transition, renewed biofacies diversification, in particular re−appearance of reef−related ancyrodellid fauna, took place. In addition, a large−scale migration event among palmatolepids and polygnathids during sea−level rise, mainly from the East European Platform, characterised this Middle Frasnian interval.
The oxygen isotopic composition of conodont apatite derived from the Late Triassic (Carnian to lower Norian), Pignola 2 and Sasso di Castalda sections in the Lagonegro Basin (Southern Apennines, Italy) was studied in order to constrain the habitat of Late Triassic conodont animals. Oxygen isotope ratios of conodonts range from 18.5 to 20.8‰ V−SMOW, which translate to palaeotemperatures ranging from 22 to 31ºC, assuming a δ18O of Triassic subtropical sea water of −0.12‰ V−SMOW. These warm temperatures, which are well comparable to those of modern subtropical−tropical oceans, along with the body features of the conodont animal suggest that conodont δ18O values reflect surface water temperatures, that the studied conodont taxa lived in near−surface waters, and that δ18O values of Late Triassic conodonts can be used for palaeoclimatic reconstructions.
In order to document the Alamo Event and to investigate its influence on shallow−marine environments, we undertook a study of ostracods, conodonts, and analysis of the sedimentology of the lower member of the type Devils Gate Limestone. Six major carbonate microfacies (MF1–MF6) ranging from open−marine environments below storm wave base to pre−evaporitic supratidal lagoons were recognized. The sedimentological study detected no important sedimentological changes during the Alamo Event; only an influx of detrital material and lithoclasts indicate that an unusual event had occurred. Ostracods are generally rare or absent in the lower member of the Devils Gate Limestone, and only 2,000 carapaces, valves and fragments were extracted; from these some 26 taxa were identified. Two new species, Voronina? eureka and Serenida dorsoplicata are proposed. The ostracods belong to the Eifelian Mega−Assemblage and their distribution was influenced by strong salinity variations. Because of the rarity and low diversity of ostracods and conodonts in samples collected from the lower part of the lower member of the Devils Gate Limestone it is not adequate to demonstrate conclusively an extinction event close to the Alamo Event Bed. Nevertheless the greater abundance and diversity of ostracods above this bed seems to indicate that the Alamo Event did not result in significant extinction of ostracod taxa in this shallow water setting. The ostracod fauna present in the lower member of the Devils Gate Limestone suggests faunal exchanges between Nevada and the Russian Platform via the Western Canadian platform.
The Frasnian–Famennian (F–F) boundary is well biostratigraphically documented in the Palmatolepis−rich deposits exposed along the Syv’yu River in the lower slopes of the Subpolar Urals. The thin−bedded calcareous−clayey−siliceous deep−slope succession of the Vorota Formation appears to represent continuous Domanic−type deposition throughout the world−wide carbonate crisis time, without evidence for the basal Famennian hiatus or a large−scale sedimentary perturbation within a regressive setting. The northernmost Laurussian sequence exhibits many well known signatures throughout the broad F–F timespan: the appearance of organicand clay−rich deposits, icriodontid and radiolarian blooms, and a correlative shift of several geochemical proxies towards hypoxic and high−productivity regimes, perfectly recorded by positive 13Ccarb excursions of +3.5‰. Integrative biotic, microfacies and geochemical data substantiate a longer−term oceanographic destabilization, attributable to multiple Earth−bound triggering factors in (episodically enhanced?) greenhouse climate and punctuated eustatic sea−level highstands, superimposed on the elevated deposition of organic carbon−rich sediments during the Upper Kellwasser Event. Unsteady eutrophicated, and oxygen−depleted ecosystems during the F–F biotic crisis interval could be assumed, especially when intensified by various spasmodic tectono−volcanic phenomena in the incipiently closing Ural Ocean.
The dominant factor in faunal succession of conodonts in the Frasnian of Poland is the apparent immigration of species originating allopatrically in other regions. Each immigration event usually changes the population variability of a local species (character displacement). Only a few lineages show their phyletic evolution within the studied area. Attempts to distinguish conodont species on the basis of platform element shape failed in some of the latest Frasnian palmatolepidids. Even at the apparatus−based generic level, certain ramiform elements of the apparatus appear much more diagnostic than the platforms. Correlative value of the late Frasnian palmatolepidids of unknown apparatus structure is thus questionable. The evolution of platform elements in Ancyrodella offers a more solid basis for age determination although their great population variability makes resolution rather low and requires the population approach. The panderodontids Belodella(?) tenuiserrata sp., B. minutidentata sp. nov., B. robustidentata sp. nov., prioniodontid Icriodus kielcensis sp. nov., enigmatic monospecific Playfordiidae fam. nov., prioniodinids Dyminodina planidentata gen. et sp. nov., D. anterodenticulata sp. nov., D. kovalensis sp. nov., Pluckidina kielcensis gen. et sp. nov., P. slupiensis sp. nov., P. robustipegmata sp. nov., and P. lagoviensis sp. nov., derived polygnathid Avignathus bifurcatus sp. nov., probably secondarily simplified polygnathid Nicollidina gen. nov., and palmatolepidids Kielcelepis gen. nov., Lagovilepis gen. nov. and Klapperilepis gen. nov. are proposed.
Scolecodonts and conodonts are described from five samples of three cores from the Svalis Dome, central Barents Sea. The conodont species Mesogondolella rosenkrantzi and Neospathodus svalbardensis confirm latest Permian (Dzulfian) and earliest Triassic (Dienerian) ages for the investigated intervals. Correlation with adjacent conodont distribution indicates a shorter depositional gap in the late Permian than is evident on Svalbard. The conodont elements show little thermal alteration with CAI (colour alteration index) values between 1.0 and 1.5 indicating a maximum burial temperature of 50—90°C. The scolecodonts described herein have important evolutionary and distribution implications for polychaetes: the order Eunicida shows a richer diversification in the Permian than previously known; the family Paulinitidae survived at least until the Late Permian; the family Hartmaniellidae diversified already in the late Palaeozoic; some forms are considered as representatives of unknown species and genera; the order Phyllodocida originated already in the latest Palaeozoic.
The Lower Ordovician conodont Scolopodus striatus Pander, 1856 (= Scolopodus rex Lindström, 1955) has an apparatus composed of five element morphotypes: acontiodiform, subrounded, compressed paltodiform, paltodiform, and scandodiform. The identification of the morphotypes is based on the general asymmetry of elements and shape of their bases. The elements are variable within each morphological group and form a continuous transition series generally reflected in differences in the depth of the basal cavity, height and degree of lateral compression of the base. The new collection from the localities near St. Petersburg, the type area of the first investigations on conodonts by Christian Pander in 1856, was examined and species of Scolopodus named by him are revised. All Pander’s species with the exception of the type species Scolopodus sublaevis are identified as a single species of Scolopodus that was named 99 years later as Scolopodus rex. The S. sublaevis sensu formae was not recognised in collections studied and its validity is questionable. S. striatus is the most easily recognisable among Pander’s species of Scolopodus and is here proposed to be a senior synonym of S. rex.
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