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Megalospheric forms of Western Tethyan late Bartonian to late Priabonian involute Heterostegina from numerous localities, marking different ecological conditions, were morphometrically investigated. They belong to three species, H. armenica, H. reticulata, and H. gracilis based on the presence/absence of granulation, on the chamberlet characteristics and on the relative size of proloculus. Within these species a very rapid evolution could be observed in the reduction of the number of operculinid chambers, in the increase of the number of chamberlets and partially in the increase of the proloculus size. This evolution is demonstrated by stratigraphic superpositions in several localities (especially in the Mossano section), and is supported also by the change of co−occurring fossils, starting with the disappearance of large−sized Nummulites, then followed by the appearance of the genus Spiroclypeus and then by the disappearance of orthophragmines of middle Eocene acme. Based on the reduction of operculinid chambers, two chronosubspecies of Heterostegina armenica and seven of H. reticulata are defined biometrically (four of them: H. armenica tigrisensis, H. reticulata tronensis, H. r. hungarica, and H. r. mossanensis are introduced here). This allows to subdivide the Shallow Benthic Zone (SBZ) 18 into three and SBZ 19 into two subzones. The extremely rapid evolution of H. reticulata allows to calibrate larger foraminiferal events around the middle/late Eocene boundary. The extinction of large−sizedNummulitesseems to be heterochronous in the late Bartonian in having migrated eastward, while the first appearance of Spiroclypeus is shown to be synchronous at the base of the Priabonian. The middle/upper Eocene (= Bartonian/Priabonian) boundary is to be placed at the base of the Priabona marls in the Mossano section corresponding to the SBZ 18/19 limit, to the first appearance of genus Spiroclypeus, to that of Nummulites fabianii and of Heterostegina reticulata mossanensis. It falls into the upper part of both the P 15 and NP 18 planktic zones. The Western Tethyan Eocene involute Heterostegina became extinct, apparently with no Oligocene successors.
Megalospheric forms of Priabonian Spiroclypeus of the Western Tethys were morphometrically investigated. Based on the reduction of the average number of undivided, post−embryonic chambers, the investigated populations are grouped into two successive, phylogenetically linked species, S. sirottii sp. nov. and S. carpaticus. The evolution is also demonstrated by the increase of the number of secondary chamberlets in particular chambers, by the increase of the diameter of the first two whorls and by that of the size of the proloculus, although the latter turned out to be also ecologically controlled. This evolution is supported by the stratigraphic succession of populations in the Mossano section (N Italy) and by the change of accompanying fossils. Lacking in upper Bartonian beds, the first appearance of genus Spiroclypeus seems to be synchronous with the beginning of the late Eocene. The newly described S. sirottii is associated with Heterostegina reticulata mossanensis and orthophragmines containing forms of middle Eocene acme, both marking the lower part of the Priabonian. Meanwhile S. carpaticus co−occurs with H. gracilis and/or with orthophragmines characteristic of the upper part of the Priabonian. We suppose that the Spiroclypeus sirottii–carpaticus lineage is restricted to the Priabonian. Thus, Spiroclypeus sirottii is a zonal marker for the Shallow Benthic Zone (SBZ) 19 (early Priabonian) while S. carpaticusindicates the SBZ 20 (late Priabonian).
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