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An early Late Tournaisian (Early Carboniferous/Mississippian) ammonoid fauna is described from the Tafilalt of south−eastern Morocco. Twelve genera, four of which are new, and eleven new species are represented: Becanites africanus sp. nov., Triimitoceras epiwocklumeriforme gen. et sp. nov., Irinoceras minutum sp. nov., Muensteroceras quadriconstrictum sp. nov., Eurites bouhamedensis sp. nov., Ouaoufilalites ouaoufilalensis gen. et sp. nov., Helicocyclus fuscus sp. nov., Pericyclus mercatorius sp. nov., Orthocyclus(?) sp., Bouhamedites enigmaticus gen. et sp. nov., Winchelloceras antiatlanteum sp. nov., and Progoniatites maghribensis gen. et sp. nov. Palaeogeographic analysis of Late Tournaisian ammonoid assemblages shows strong endemism at the species−level, but genera and families had a nearly global distribution in the equatorial seas. The new fauna contains the stratigraphically oldest known representatives of the important Carboniferous goniatite families Girtyoceratidae and Goniatitidae.
The fossil record of Paleozoic ophiuroids includes a number of forms which share striking similarities with modern relatives in terms of skeletal morphology. These so called modern-type Paleozoic ophiuroids yield an enormous potential for a better understanding of ophiuroid evolution, yet the scarcity of accurate and sufficiently detailed morphological descriptions available to date precludes any further-reaching assessments. Here, we describe an articulated ophiuroid specimen from the Late Tournaisian (early Carboniferous) of Czatkowice quarry, southern Poland, as a new species Aganaster jagiellonicus sp. nov. The good preservation of the specimen allowed for a morphological analysis at a level comparable to recent ophiuroid descriptions. It shows remarkable morphological similarities with extant former ophio-lepidids Ophiomusium and Ophiosphalma. The new find thus contributes to a solid basis for future investigations on the position of the modern-type Paleozoic ophiuroid in the phylogeny of the class.
The shallow water assemblage of chondrichthyan microremains, teeth, tooth plates and scales, from the middle Tournaisian (Mississippian) of the vicinity of Muhua village, Guizhou province, southern China, is thus far the richest and most diverse association of this age collected from a single locality and horizon, and represents a chondrichthyan community very restricted in time and space. It was recovered from a small bioclastic limestone lens, MH−1, occurring among basinal marls near the base of the Muhua Formation, and dated as to the Siphonodella crenulata conodont Zone. The majority of the fauna presented here consists of teeth with euselachian−type bases and crushing crowns belonging to bottom−dwelling durophagous chondrichthyans, most probably feeding on shelly invertebrates such as the abundant brachiopods. We assigned most of these teeth to Euselachii (six species, among them Cassisodus margaritae gen. et sp. nov.), Petalodontiformes (two species), Holocephali (five species), and Euchondrocephali incertae sedis (Cristatodens sigmoidalis gen. et sp. nov.). We also identified primitive polycuspid, clutching teeth representing Phoebodontiformes (Thrinacodus bicuspidatus sp. nov.), Symmoriiformes, and Ctenacanthiformes. The scales are typical growing, compound forms of the protacrodont, ctenacanth, and hybodont types. Two problematic denticulated plates were found, one of which resembles mandibular or palatal plates of Sibyrhynchus (Iniopterygii). Several of the identified chondrichthyan taxa have hitherto been known only from Laurussia, especially from the British Isles and central USA. In particular we found the first record of Chondrenchelyssp. and Diclitodus denshumani outside their type locality. Th. bicuspidatus sp. nov., also known from Nevada, Iran, and NW Australia, appears to be a cosmopolitan, middle Tournaisian index fossil.
An uppermost Famennian (Strunian) coral assemblage has been recovered in the middle part of the Yılanlı Formation of the Istanbul Zone (Zonguldak and Bartın areas, NW Turkey). In the Bartın area, the studied fossiliferous interval corresponds to a c. 30 m-thick unit of bioclastic to peloidal wackestone to packstone grading to grainstone and including two stromatoporoid biostromes. In the Zonguldak area, 60 km westward, the bioclastic facies is dominant. The rugose corals are mainly solitary taxa belonging to the genera Campophyllum, Bounophyllum, Amplexocarinia, and ?Metriophyllum, and only one colonial genus occurs: Pseudoendophyllum. This fauna is similar to that documented in Europe. The campophyllids and dibunophyllids are the main component of the uppermost Famennian assemblages in S Belgium, N France, W Germany, NW and S Poland. The endophyllids occur in S Poland, Novaya Zemlya, and in the Ural Mountains. The Istanbul Zone is supposed to be situated in the central part of the Palaeotethys Ocean, along the southern margin of Laurussia during the uppermost Devonian and Carboniferous. The rugose corals indicate some relationship with the eastern part of Laurussia, or that both areas were under a common marine influence at this time. The global Hangenberg event was not recognized in the Turkish localities, except for the disappearance of the corals, occurring less than 19 m below the Devonian–Carboniferous boundary based on the foraminifers. There is no major facies change through the boundary and the first Carboniferous corals (small Uralinia and Caninophyllum) appear 6 m above the D–C boundary. The new species Caninophyllum charli sp. nov. is described from the upper part of the lower Tournaisian.
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