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A new genus of middle Tremadocian orthoceratoids and the Early Ordovician origin of orthoceratoid cephalopods

100%

Kroger B.

Acta Palaeontologica Polonica

2008

tom 53

nr 4

The cephalopods of the subclass Orthoceratoidea, which are termed “orthoceratoids” herein, are a group that remains “the last unexplored wilderness in the Cephalopoda” (Flower 1962: 23). After 45 years this statement still holds true because phylogeny reconstructions are hindered by their morphological simplicity, numerous homeomorphies and iterative evolution. The Orthocerida, straight cephalopods that are characterised by a wide chamber spacing, a thin tubular siphuncle and a small spherical initial chamber, lacking a cicatrix (Kröger 2006) were the ancestors of bactritoids, ammonoids, and coleoids (Engeser 1996). The origin of the Orthocerida is poorly understood. The earliest unequivocal Orthocerida are known from the Floian (Early Ordovician). A number of poorly known possible Orthocerida and/or stem group Orthocerida are known from the Tremadocian. Here, I reassign the long known middle Tremadocian “Orthoceras attavus” to the new genus Slemmestadoceras belonging to a group of worldwide distributed orthoceratoids. The presence of Slemmestadoceras with a thin, probable tubular siphuncle and small initial chambers in the middle Tremadocian suggests that the Orthocerida may have originated already at that time. The comparison of Slemmestadoceras with following late Tremadocian and Floian orthoceratoids demonstrates that a higher level taxon comprising these forms, such as the subclass Orthoceratoidea may constitute a paraphylum.

Soft-tissue attachments in orthocerid and bactritid cephalopods from the Early and Middle Devonian of Germany and Morocco

80%

Kroger B., Klug C., Mapes R.

Acta Palaeontologica Polonica

2005

tom 50

nr 2

In the Early to Middle Devonian shale sequences of Germany and Morocco, pyritised and secondarily limonitised cephalopod remains are common. Details of the soft−tissue attachment structures are sometimes preserved on the internal moulds of the body chamber and phragmocone of these cephalopods. Some of the studied Orthocerida show a very faint annular elevation and a dorsal furrow. A few Bactritida show a distinctive annular elevation with two circular bands. The bands form a paired or threefold lobe at the dorsum of the shell. Morphological differences between Orthocerida and Bactritida suggest different soft part morphologies. A comparison of the attachment scars shows that the Bactritida are intermediate between the Orthocerida and ammonoids with regard to their muscle attachment scars. The shape of the muscle scars are interpreted as indication for a planktonic lifestyle in Orthocerida and a comparatively active, nektonic lifestyle in Bactritida and ammonoids. The new genus Acanthomichelinoceras is erected. Acanthomichelinoceras commutatum, Cycloceras sp., Bactrites gracile, Bactrites sp. A, Bactrites sp. B, and Bactrites sp. C are described.

Ograniczanie wyników

2 Acta Palaeontologica Polonica

2 Kroger B.

1 Klug C.

1 Mapes R.

1 2008

1 2005

Wyniki wyszukiwania

A new genus of middle Tremadocian orthoceratoids and the Early Ordovician origin of orthoceratoid cephalopods

Soft-tissue attachments in orthocerid and bactritid cephalopods from the Early and Middle Devonian of Germany and Morocco