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Late Frasnian Atrypida (Brachiopoda) from the South Urals, South Timan and Kuznetsk Basin in Russia (east Laurussian and south Siberian shelf domains in Devonian time) reveal significant generic and specific diversity in the broadly defined Frasnian-Famennian (F-F) bio-crisis time. Eighteen species of atrypid brachiopods have been recorded, representing 4 subfamilies and 10 genera. The new genus Gibberosatrypa Markovskii & Rzhonsnitskaya, and the new subgenus Spinatrypa (Plicspinatrypa) Rzhonsnitskaya are proposed. Four new species Spinatrypina (Spinatrypina) sosnovkiensis Yudina, Spinatrypa (Plicspinatrypa) rossica Rzhonsnitskaya, Iowatrypa nalivkini Rzhonsnitskaya & Sokiran, and Cartnatina(?) biohermica Yudina are described. The representatives of the Variatrypinae (including especially common Desquamatia (Desquamatia) alticoliformis), Spinatrypinae (Spinatrypina) and Atypinae (Pseudoatrypa, ?Costatrypa) are widely distributed in the studied regions. The Pseudogruenewaldtiinae are represented by Iowatrypa and Pseudogruenewaldtia, of which the first is distributed worldwide, whereas the only undoubted species of the second is restricted to South Timan, and probably represents a localized latest Frasnian descendant of Iowatrypa. The decline phase of atrypid development was controlled by a variety of environmental factors tied to the global Kellwasser events, although it was not directly triggered by anoxic conditions. The investigated atrypid brachiopods, which were all confined to lower latitudes, disappeared during the F-F mass extinction, independently of their environmental and biogeographic settings.
The brachiopod fauna of the Middle-Late Devonian cratonic carbonate platform deposits of the Iowa Basin, central North America, contains twenty species of the order Atrypida, some of which are types for widespread genera common in Middle and Late Devonian faunas. The latest Givetian-early Frasnian deposits yield a diverse fauna consisting of ten species (two new) included in Desquamatia (Independatrypa), D. (Seratrypa), Pseudoatrypa, Radiatrypa, Spinatrypa (Spinatrypa) and Spinatrypina (Exatrypa). Many of these forms occur in, or are closely similar to species known from, coeval faunas of central and western Canada. Middle Frasnian deposits of northern Iowa contain two species included in Spinatrypa (S.) and Pseudoatrypa, both of which are new. Late Frasnian strata of the Iowa Basin yield eight species included in Costatrypa, Iowatrypa, Pseudoatrypa, and Spinatrypa (Spinatrypa), some of which are widespread in other subtropical and tropical faunas of the western US and western Canada. The taxa Pseudoatrypa witzkei sp. n., Spinatrypa (S.) bunkeri sp. n., Spinatrypa (S.) thompsoni sp. n., and Spinatrypina (Exatrypa) johnsoni sp. n. are proposed. Pseudoatrypa? sp. from the very late Frasnian of southern New Mexico is also illustrated.
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