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The species richness, H’ and D’ diversity and evenness J in southwestern- and northeastern-facing slopes were compared and height of twelve herbaceous plant species was measured five times during vegetation season in old-growth beech forest. The opposite slopes chosen for studies were similar in relation to slope inclination, type of soil, species composition and canopy cover. Two study plots 60 m2 encompassing two rows of 1- by 2-m 15 rectangular quadrates2 were established. The study revealed that northern slopes were characterized by significantly lower mean species diversity (H’ = 0.82; D’ = 0.47) compared to southern slopes (H’ = 1.2; D’ = 0.59), similar evenness (ca. J = 0.72) and higher heterogeneity with respect to spatial structure using Euclidean distance and range of DCA scores of quadrates of plots. Two-way ANOVA yielded significant differences of effect of slope aspect (P <0.001) and period of time (P <0.01) on average vegetation height. Nevertheless, interaction of time and aspect turned out to be insignificant. The highest mean values of height of plants were observed at the end of July and the end of June. Generally plants growing on north-facing slope were taller and had higher cover than those growing in southern sloped forest. This can be the result of lower species diversity on northeastern slope which favours development of individual species in conditions of weak interspecific competition and stronger intraspecific interactions.
The paper presents results of the research carried out in beech stands of the Carpathian beech fertile complex (Dentario glandulosae-Fagetum) in the lower subalpine zone in the Bieszczady National Park. The objective of the presented study was to analyse changes of species composition in all developmental stand of the lowers stands in the period from 1993 to 2003/04. The obtained results are proof of high levels of stability of the stands in focus, which was confirmed by absence, at the beginning and at the end of the control period, of statistically significant changes in species composition of the old trees layer, undergrowth and natural seeding. Enduringly, the Carpathian beech stands are build by three specific forest-forming species to the Eastern Carpathian Mts.: beech, fir and sycamore.
The paper presents the results of studies on bryophyte flora of the Reberce forest reserve located in the Przemyśl foothills. Thirty-three species of liverworts, 70 species and two varieties of mosses were found. Terrestrial (66 species) and epixylic (58) bryophytes formed the largest ecological groups in the reserve. Twenty-four protected species were also recorded, eight endangered bryophytes and six taxa considered as primeval forest species. The most interesting finds are: Bazzania trilobata, Buxbaumia viridis, Calypogeia suecica, Cephalozia catenulata, Frullania dilatata, Metzgeria conjugata, Nowellia curvifolia, Orthotrichum lyellii, O. stramineum, Trichocolea tomentella and Ulota crispa.
From among three stands, situated in the lower mountain zone, and representing the association Dentario-glandulosae Fagetum, the Łopuszna I stand, being in the growing up stage and phase of selection forest, reached the highest increment (8.5 m3/ha/year, i.e. 1.3% of actual stand volume/ha measured at the end of the control period). Stands, Łopuszna II, being in the stage of an intensive break-up, and Łopuszna III, being in the stage of an initial break-up, reached smaller increments (5.4 and 6.3 m3/ha/year respectively, i.e. each 0.9% of actual stand volume/ha). The greatest tree mortality occurred in stands Łopuszna II and III (14.1 and 10.5 m3/ha/year respectively), and the smallest in the stand Łopuszna I (4.8 m3/ha/year). The knowledge about the value of mortality, recruitment, and increment, expressed in the number of trees and in volume units, may greatly help in planning the amount of cut which would secure sustainability of the forest ecosystem in the layer of trees in the economic as well as in the protection forests, under similar stand and site conditions, managed by the selection cutting permitting to realize the conception of a close-to-nature silviculture.
Long-term changes in groundlayer vegetation of unique stand of Carpathian beech forest Dentario glandulosae-Fagetum considered partly as a natural succession due to climatic changes and partly as a response to canopy dynamics, were investigated in the Ojcow National Park (South Poland). Species composition and structure of the herb layer in the permanent research area were compared in 1993 and after ten years. About 77% of the study area was changed in this period, but the spatial pattern of distribution and intensity was highly mosaic. The mean intensity of changes calculated according to formula (1) derived from changes in percent cover of species was 42%. Herb cover decreased by about 15%, probably due to the rapid development of natural recruitment of trees in the study area (48% of area was covered by dense tree ecruitment). In the same time the number of plant species and vegetation types (homogeneous vegetation patches of the size at least 1m2, that differed from neighbouring patches in species composition and/or species share) increased substantially. It is probably due to greater variation of light condition in ground layer created as a result of canopy gaps dynamics (the area of windthrow mounds in the study area increased approximately eight times between 1993 and 2003). There was a significant decline of species typical for beech forests, especially of vernal geophytes like Dentaria glandulosa W.K. and Isopyrum thalictroides L. Although it is too early to predict any directional changes, it is possible that the beech forest community will change into the Tilio-Carpinetum association in some parts of study area. Nowadays the expansion of invasive alien species Impatiens parviflora DC. is the great threat to the forest vegetation of the protected study area and the Ojcow National Park. This species in 10 years managed to displace native species in 3% of the study area. The drastic decrease in herbaceous plants cover connected with increase in tree seedlings and saplings, systematic decline of vernal geophytes and typical beechwood species, diversification of ground vegetation structure and composition, and finally occurrence of non-native invasive plant species inform about highly dynamic nature of climax beech forests and generally of temperate forests. Changes are induced probably by both changing climate and overstorey dynamics.
The structure of carabid assemblages was studied in the Holy Cross Mountains, in stands of Luzulo pilosae-Fagetum lowland acidic beech forest (LAB), Dentario glandulosae-Fagetum fertile Carpathian beech forest (FCB), and Dentario enneaphylli-Fagetum fertile Sudeten beech forest (FSB) in 2002. Carabid beetles were collected using Barber's pitfall traps with ethylene glycol in monthly cycles from May to September. A total of 4260 individuals representing 28 species were caught. The highest similarity was calculated for the LAB and FCB assemblages (P=0.54), while the FCB and FSB assemblages were the least similar (P=0.39). The highest diversity was recorded in the FSB assemblage (H'=2.06), and the lowest in the assemblage of acidic lowland beech stands (LAB) (H'=1.45). The dominant species were Carabus violaceus in the LAB sites, Carabus glabratus in the FCB sites, and Carabus hortensis in the FSB sites. The most uniform distribution of individuals between species was seen in the FSB assemblage (0.76) and in the FCB assemblage (0.74), while the LAB assemblage was highly non-uniform in this respect (0.53) Trapability was lowest in LAB (0.36) and highest is FSB (0.55). The prevailing zoogeographical elements were Palaearctic species in LAB, European Forest Province montane species in FCB, and European Forest Province species in FSB. The LAB and FCB assemblages were most active in the spring (Fig. 4) and the FSB assemblage reached peak activity in the summer (July). The dominant species types were forest species (habitat preferences), large zoophages (feeding preferences), mesohygrophiles (humidity preferences), and spring breeders (developmental type).
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