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The Gobi Desert is famous for providing one of the worlds best preserved Cretaceous terrestrial faunas, including dinosaurs and mammals. Beginning with the Central Asiatic Expeditions in the 1920s, through the Polish−Mongolian Expeditions in the 1960s–1970s, Soviet−Mongolian Expeditions in 1970s, and finally the Mongolian Academy−American Museum Expeditions in the 1990s–2000s, the number of complete skulls (see Kielan−Jaworowska et al. 2000 for review) of Cretaceous mammals often associated with postcranial skeletons, found in Mongolia increased to several hundred. In addition to these professional expeditions, there have been other types of trips to Mongolia, also aimed at collecting fossils. The Nomadic Expeditions Company in USA organizes one of these, and has made trips to Mongolia since 1996. During the 1999 Nomadic Expedition, a skull associated with parts of the postcranial skeleton of the multituberculate mammal Catopsbaatar catopsaloides was found. The specimen is more complete than others previously known of this species and brings new data on multituberculate anatomy and ontogenetic variation. In this note we discuss the new data on the structure of C. catopsaloides; the details of its anatomy will be described in subsequent papers by the two first authors.
We describe an incomplete postcranial skeleton of Catopsbaatar catopsaloides from the ?late Campanian red beds of Hermiin Tsav I, in the Gobi Desert, Mongolia. The skeleton is fragmentary and the preservation of bone surface does not permit reconstruction of the musculature. The studied skeleton contains some parts not preserved or incompletely known in other multituberculate genera, such as a long spinous process in a single lumbar vertebra, which together with long transverse processes preserved in Nemegtbaatar, might indicate that at least some multituberculates had jumping ability. The calcaneus of Catopsbaatar is unusual, differing from most other multituberculates (where known) and other mammals by having a short tuber calcanei, with a large proximal anvil−shaped process strongly bent laterally and ventrally, arranged obliquely with respect to the distal margin of the calcaneus, rather than arranged at 90° to it, as in other mammals. This suggests the presence of strong muscles that attached to the tuber calcanei, perhaps further attesting to jumping abilities in Catopsbaatar. We also describe an unfused pelvic girdle and the first extratarsal spur bone (os cornu calcaris) known in multituberculates.
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Mongolian Late Cretaceous multituberculates (except Buginbaatar) form a monophyletic group for which the suborder Djadochtatheria is proposed. Synapomorphies of Djadochtatheria are: large frontals pointed anteriorly and deeply inserted between the nasals. U-shaped fronto-parietal suture, no frontal-maxilla contact, and edge between palatal and lateral walls of premaxilla. Large, rectangular facial surface of the lacrimal exposed on the dorsal side of the cranial roof is present in all djadochtatherians, but may be a plesiomorphic feature. It is also possible that in djadochtatherians the postglenoid part of the braincase is relatively longer than in other multituberculates. Djadochtatherians have an arcuate p4 (secondarily subtrapezoidal in Catopsbaatar) that does not protrude dorsally over the level of the molars (shared with Eucosmodontidae), I3 placed on the palatal part of the premaxilla (shared with the eucosmodontid Stygimys and the cimolomyid Meniscoessus). The small number of cusps on the upper and lower molars and no more than nine ridges on p4 are possibly plesiomorphies for Djadochtatheria. The djadochtatherian Nessovbaatar multicostatus gen. et sp. n., family incertae sedis from the Barun Goyot Formation is proposed. New specimens of the djadochtatherian genera Kryptobaatar, ?Djadochtatherium, and Kamptobaatar are described and revised diagnoses of these taxa and Sloanbaatar are given. A cladistic analysis of Mongolian Late Cretaceous multituberculates (MLCM), using Pee-Wee and NONA programs and employing 43 dental and cranial characters, 11 MLCM taxa, five selected Late Cretaceous or Paleocene multituberculate genera from other regions, and a hypothetical ancestor based on the structure of Plagiaulacoidea, is performed. The Pee-Wee program yielded two equally fit trees that confirm the monophyly of MLCM excluding Buginbaatar. Kryptobaatar, Djadochtatherium, Catopsbaatar, and Tombaatar form a clade, for which the family Djadochtatheriidae is proposed. Chulsanbaataris the sister taxon of this clade. Bulganbaatar and Nemegtbaatar are the sister group of all other djadochtatherians. Kamptobaatar, Sloanbaatar, and Nessovbaatar form a separate clade in the Pee-Wee tree. The NONAprogram yielded thirty equally parsimonious trees and a strict consensus tree with a poor resolution.
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