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This paper presents results of analysis of the dynamics of growth and development of beech stands in the lower subalpine zone in the Bieszczady National Park from the point of view of size-volume interrelationships of the following three main forest-forming processes: thickness increment, the process of trees dying off in stands and growing of young trees to reach the layer of the manure stand. The obtained results confirmed that the examined beech forests exhibit similarity with multigenerational, complex forests of primeval character. An argument in favour of this includes in particular: high stand volume, good forest health condition and a positive relationship between the process of tree survival (volume increment and ingrowth) and the decrement process. The wide peak distance of culminations of two traits is characteristic too: the number of trees in the lowest diameter subclass and the volume in the diameter subclass several intervals higher.
Directions and the scope of changes in the species composition of individual layers in secondary pine communities belonging to the class Querco-Fagetea Br.-Bl. & Vlieg. 1937 (tree stand, shrubs and vascular plants of the herb layer) observed at two study plots over almost 40 years (from 1971 until 2009) are discussed. Both tree stands were planted and they are in the phase of spontaneous regeneration towards a potential natural community – the Carpathian beech forest Dentario glandulosae-Fagetum Oberd. 1953. Species with greater light requirements (pine Pinus sylvestris L., aspen Populus tremula L., oak Quercus robur L., ecologically alien to the potential plant community, retreat from the tree and shrub layers. The share of tree species typical of the potential community (beech Fagus sylvatica L., hornbeam Carpinus betulus L., sycamore Acer pseudoplatanus L.) – in the tree stands increases. The rate of changes is particularly high for beech, while regression symptoms are noticed for fir Abies alba Mill.: a decrease in the abundance and share of fir in both tree-stands was recorded. A decrease in the abundance and frequency of the majority of species, including species characteristic of deciduous forests (class Querco-Fagetea) and beech forests (order Fagetalia sylvaticae Pawł.1928), and an increase in species typical of alder-ash riparian forests (alliance Alno-Ulmion Br.-Bl. & Tx. 1943) and nitrophilous communities were observed in the herb layer. Conversion treatment may reinforce the artificial character of the community because the species composition, as well as the spatial and age structure of the tree stands, can be arbitrary and therefore artificially formed during conversion (arbitrary applied size and order of regeneration clumps and areas). When anthropogenic communities are left undisturbed, processes affecting all phytocoenotic layers are activated and spontaneous forest regeneration usually begins. Therefore, tree-stand conversion used as a tool to restore natural community is not effective.
13 year changes in the ground vegetation of natural beech forest were studied in the Babia Góra National Park (South Poland) and compared with the results of similar study in the Ojców National Park published previously (Łysik 2008). In 1-ha permanent research plot homogeneous vegetation patches of the size at least 1m², that differed from neighbouring patches in species composition and/or species share (patch types) were specified and mapped. After intersecting two (for years 1991 and 2004) numerical maps a synthetic map, used then as a basis for evaluating the ground vegetation changes, was created. Changes in each patch and dynamics of each species were derived using adequate formulas. Shannon-Wiener and evenness indices were measured for both years. Comparison of species composition and structure of the herb layer in 1991 and 2004 gives evidence of a highly dynamic nature of the ground layer. On 27% of investigated area vegetation were totally different than 13 years before, while less than 3% of the study area stayed unchanged. On the rest of the area changes of diverse intensity occurred. However, the mean over-plot intensity of changes was about 65% (calculated according to formula 1). Plant cover decreased by about 20%, but at the same time increases were noticed in species number and patch types. There was a significant increase in the area covered by grasses and ferns at the expense of Rubus idaeus L., which used to be a dominant species in 1991. Moreover, cover share of all species become more evenly distributed in comparison with situation 13 years before (evenness index increased from 0.49 to 0.68). The most dynamic species were those associated with decaying stumps and downed logs. These results suggest that, in natural beech forests, overstory dynamics or other mechanisms (like competition with natural recruitment of trees) make the dominance of any species in ground-layer not longer than a decade.
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