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Earliest eutherian mammal skull, from the Late Cretaceous [Coniacian] of Uzbekistan

100%

McKenna M. C., Kielan-Jaworowska Z., Meng J.

Acta Palaeontologica Polonica

2000

tom 45

nr 1

1-54

We describe a partially crushed skull and dentaries of a sub-adult individual of Daulestes nessovi sp. n., from the Coniacian of Uzbekistan. This is the earliest known eutherian skull (about 87 Ma) and the sixth genus of Cretaceous eutherians in which a skull is available. Because the skull of D. nessovi is sub-adult, certain plesiomorphic features may be ontogenetic and should be interpreted with caution. Four upper premolars and five lower premolariform teeth were in use (possibly to become four lowers when fully adult). The upper cheek-teeth have winged conules; M2 has large parastylar and small metastylar projections. Pre- and postcingula are lacking on DP4 and the upper molars. The talonids of dp4-m2 are about 90% as wide as the trigonids, with widely separated entoconid and hypoconulid. The skull has a large sphenorbital fissure, no foramen rotundum, and apparently no pterygoid process of the sphenoid. A large orbital wing of the palatine prevents maxilla-frontal contact within the orbit. The zygomatic arch is slender. The cochlea has one full turn, with an expanded apex, which suggests that a lagena might have been present. A large malleus with a robust anterior process, and a large promontorium may be due to young age of the individual or a primitive retention, as in the platypus. Because of the similarity to Asioryctidaem both cranial structure and dentition, we assign Daulestes tentatively to Asioryctitheria Novacek et al. 1997, family incertae sedis.

Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

80%

Archibald J. D., Averianov A. O.

Acta Palaeontologica Polonica

2006

tom 51

nr 2

Four small asioryctitheres at Dzharakuduk (Turonian), Uzbekistan are Daulestes kulbeckensis (= Kumlestes olzha), D. inobservabilis (= Kennalestes? uzbekistanensis), Uchkudukodon (gen. nov.) nessovi and Bulaklestes kezbe. Uchkudukodon nessovi is one of the smallest therians (molars about 1 mm long). Lower canine is two−rooted in Uchkudukodon gen. nov. and Bulaklestes(uncertain in Daulestes). All lower premolars in all four species are double−rooted. Teeth identified as dp1, p2 and dp2 in holotype of Uchkudukodon nessovi (McKenna et al. 2000) are here identified c, p1, and p2. A phylogenetic analysis weakly supported Asioryctitheria by four synapomorphies: conular basins become distinct, the number of roots reverts to two on the lower canine, the p5 becomes longer than p4, and the metaconid on p5 is reduced and lost. Other characters diagnostic of asioryctitheres are four upper and lower premolars (arguably five upper premolars in juvenile Kennalestes), P4 has a protocone swelling or protocone, some asymmetry of the stylar shelf on M1–2, the paraconule on M1–3 is distinctly closer to the protocone than is the metaconule, protocone is of moderate height on M1–3 (70–80% of paracone or metacone height), Meckel’s groove is absent, and the mandibular foramen opens into a smaller depression on lingual side of mandibular ascending ramus. Asioryctes and Ukhaatherium are placed in Asioryctinae and along with Kennalestes are placed in Asioryctidae. Kennalestidae Kielan−Jaworowska, 1981 is a junior subjective synonym for Asioryctidae Kielan−Jaworowska, 1981. Because of uncertainties in the analysis, the positions of Daulestes, Uchkudukodon gen. nov., and Bulaklestes cannot be determined beyond referral to Asioryctitheria.

Ograniczanie wyników

2 Acta Palaeontologica Polonica

1 Archibald J. D.

1 Averianov A. O.

1 Kielan-Jaworowska Z.

1 McKenna M. C.

1 Meng J.

1 2006

1 2000

Wyniki wyszukiwania

Earliest eutherian mammal skull, from the Late Cretaceous [Coniacian] of Uzbekistan

Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria