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The size and morphological characteristics of a skull of an arctomorph carnivoran mammal from Mouillac (old collection of the Phosphorites du Quercy, of unknown age) in France closely match those of the holotype of the earliest known procyonid Pseudobassaris riggsi and another skull referred to this species, both from old collections of the Phosphorites du Quercy (Caylus and Mouillac), probably earliest Late Oligocene in age. The skull is more primitive in morphology than those of Pseudobassaris riggsi and every other known procyonid, approaching a hypothetical primitive procyonid morphotype. The only, but methodologically fundamental, departure from this morphotype is the lack of the procyonid suprameatal fossa, which is the crucial synapomorphy of the family Procyonidae. To explain the phylogenetic and taxonomic status of the arctomorph represented by the skull, three competing hypotheses are put forward. Hypothesis A, which considers the arctomorph as an individual of Pseudobassaris riggsi, assumes that the procyonid suprameatal fossa first appeared in a common ancestor of Pseudobassaris and other procyonids but was still of variable occurence within Pseudobassaris riggsi. Hypothesis B, which proposes the arctomorph as a member of a new Pseudobassaris species ancestral to Pseudobassaris riggsi, concludes that the procyonid suprameatal fossa arose in Pseudobassaris riggsi and in the Procyonidae independently, excluding Pseudobassaris from the procyonids. Hypothesis C, which recognizes the arctomorph as a representative of a new species of a new genus of the paraphyletic procyonid stem group, presumes that the procyonid suprameatal fossa originated in a common ancestor of Pseudobassaris and other procyonids after the new genus had become detached from the ancestral stock of the Procyonidae.
A phylogenetic analysis of the Oligocene and some Neogene European arctoid carnivorans of the order-group taxon Mustelida is performed, based on characters of the skull and dentition. The following classification of the revised genera is proposed: Simocyon (Ailurus, Amphictis, Bavarictis, Potamotherium (Pseudobassaris (Angustictis gen. n., Broiliana)) (Mustelictis ((Franconictis gen. n., Stromerielta) (Bathygale gen. n. (Plesictis (Paragale, Plesiogale)))))). Potamotherium. is allied to phocida within the monophyletic Pinnipedia. Pseudobassaris, Angustictis gen. n., and Broiliana are con­sidered procyonids. Mustelictis, Franconictis gen. n., StromerieUa, Bathygale gen. n., Plesictis, and the mustelines Paragale and Plesiogale are placed in the Mustelidae. Parailurus is included in Ailurus, and Ichneugale (= Viretius, = Aiopecocyon) is syno- nymized with Amphictis. Phylogenetic definitions and diagnoses are provided for the suprageneric taxa: Carnivora, Caniformia, Arctoidea, Arctomorpha (new), Mustelida, Pinnipedia, Procyonidae, Mustelidae, and Mustelinae.
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