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The anterior part of a medium-sized shark surrounded by hundreds of ostracod shells was found at the end of the last century in a dark limestone nodule from the Kinderscoutian (Bashkirian, Pennsylvanian) near Carsington, Derbyshire (England, UK). The shark is a caseodontoid eugeneodontiform, most probably belonging to Campodus agassizianus. Its dentition is of the crushing type, highly heterodont, but the symphyseal tooth whorl, typical of the most of eugeneodontiforms, was not found. The teeth are symmetrical labio-lingually which is unique in this group. The analysis of the available data leads to the conclusion that neither the tooth whorls of “Campodus variabilis” sensu Eastman, nor the mandibular dentition of “Agassizodus variabilis” sensu St. John and Worthen represent the genus Campodus and that these specimens deserve a new, probably common, name.
New chondrichthyan microremains from several Frasnian–Famennian sections in the Holy Cross Mountains and Dębnik area (Southern Poland) are investigated and compared to previous data. The reaction of different groups of chondrichthyans to environmental changes during the Kellwasser Event is analysed. Following the extinction of phoebodont sharks of Phoebodus bifurcatus group before the end of the Frasnian, only two chondrichthyan species, viz. Protacrodus vetustus Jaekel, 1921 and Stethacanthus resistens sp. nov. (possibly closely related to “Cladodus” wildungensis Jaekel, 1921), occur in the upper part of Frasnian Palmatolepis linguiformis conodont Zone and persist into the Famennian. Global cooling is considered a possible cause of the extinction of Frasnian subtropical phoebodonts on Laurussian margins.
The revision of shark teeth from the Pennsylvanian black shales of central USA, ascribed to “Phoebodus heslerorum” Williams, 1985 and Symmorium reniforme Cope, 1893, shows that “Ph. heslerorum” is a junior synonym of “Cladodus” divergens Trautschold, 1879. This species belongs neither to Phoebodus nor to Cladodus, so a new genus Heslerodus is proposed. Very common, robust cladodont teeth with a deep labio−basal depression and two buttons, often referred to as S. reniforme, do not belong to the latter species, but to “Cladodus” occidentalis Leidy, 1859. The generic affinity of “C.” occidentalis is yet undetermined, but it is possible that it represents ctenacanthoids.
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Devonian phoebodont shark teeth

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Shark teeth of the phoebodont type are the most common and diverse group of Upper Devonian ichthyoliths in the pelagic facies of the Holy Cross Mountains (Poland), South Urals and Timan (Russia). They were also found in the Givetian of Kuznetsk Basin (western Siberia). The morphology and function of tooth apparatus of Phoebodus was possibly similar to that of the recent shark Chlamydoselachus anguineus. A significant loss of diversity and relative productivity has been observed among the phoebodonts in the earliest Famennian. A new genus, Omalodus gen. n., and three new species of Phoebodus, Ph. bifurcatus sp. n., Ph. fastigatus sp. n. and Ph. turnerae sp. n. are proposed.
Two new assemblages of Mississippian pelagic chondrichthyan microremains were recovered from the pelagic limestone of the area of Krzeszowice, NW of Kraków, Poland. The older assemblage represents the upper Tournaisian of Czatkowice Quarry and the younger one the upper Viséan of the Czernka stream valley at Czerna. The teeth of symmoriiform Falcatidae are the major component of both collections. A comparison of the taxonomic composition of the assemblage from Czerna (with the falcatids and Thrinacodus as the major components) to the previously published materials from the Holy Cross Mountains (Poland), Muhua (southern China), and Grand Canyon (Northern Arizona, USA) revealed the closest similarity to the first of these, probably deposited in a deeper water environment, relatively far from submarine carbonate platforms. A short review of Mississippian falcatids shows that the late Viséan–Serpukhovian period was the time of the greatest diversity of this group.
The shallow water assemblage of chondrichthyan microremains, teeth, tooth plates and scales, from the middle Tournaisian (Mississippian) of the vicinity of Muhua village, Guizhou province, southern China, is thus far the richest and most diverse association of this age collected from a single locality and horizon, and represents a chondrichthyan community very restricted in time and space. It was recovered from a small bioclastic limestone lens, MH−1, occurring among basinal marls near the base of the Muhua Formation, and dated as to the Siphonodella crenulata conodont Zone. The majority of the fauna presented here consists of teeth with euselachian−type bases and crushing crowns belonging to bottom−dwelling durophagous chondrichthyans, most probably feeding on shelly invertebrates such as the abundant brachiopods. We assigned most of these teeth to Euselachii (six species, among them Cassisodus margaritae gen. et sp. nov.), Petalodontiformes (two species), Holocephali (five species), and Euchondrocephali incertae sedis (Cristatodens sigmoidalis gen. et sp. nov.). We also identified primitive polycuspid, clutching teeth representing Phoebodontiformes (Thrinacodus bicuspidatus sp. nov.), Symmoriiformes, and Ctenacanthiformes. The scales are typical growing, compound forms of the protacrodont, ctenacanth, and hybodont types. Two problematic denticulated plates were found, one of which resembles mandibular or palatal plates of Sibyrhynchus (Iniopterygii). Several of the identified chondrichthyan taxa have hitherto been known only from Laurussia, especially from the British Isles and central USA. In particular we found the first record of Chondrenchelyssp. and Diclitodus denshumani outside their type locality. Th. bicuspidatus sp. nov., also known from Nevada, Iran, and NW Australia, appears to be a cosmopolitan, middle Tournaisian index fossil.
Taxonomy of the Late Devonian placoderm remains from the Holy Cross Mountains, Poland, described by Gorizdro-Kulczycka (1934, 1950) and Kulczycki (1956, 1957), is revised. Several recently found specimens are also mentioned. The old collections are composed of representatives of Ptyctodontidae, Holonematidae, Plourdosteidae, Pholidosteidae, Selenosteidae, Titanichthyidae and Dinichthyidae, the latter with an undescribed species of Eastmanosteus. Newly found specimens belong to Ptyctodontidae, Plourdosteidae and Dinichthyidae. The occurrence of the Antiarcha in the Late Devonian of the Holy Cross Mountains, suggested by former authors, has not been confirmed.
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The first Devonian holocephalian tooth from Poland

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A recently found “bradyodont” holocephalian tooth from bituminous shales of the Kowala Quarry, south−western Holy Cross Mountains, Poland, dated as the middle Famennian Palmatolepis trachytera conodont Zone, is described. In spite of its resemblance to the forms often attributed to Helodus, the tooth is referred to as Psephodus cf. magnus (Agassiz, 1838), and supposed to represent the anterior part of the dentition, based on a partly articulated specimen of Psephodus from the Carboniferous of Scotland. The analysis of early helodonts and psephodonts, and other Famennian chondrichthyan crushing teeth, shows numerous similarities in tooth−base structure, such as the reduction of lingual basal extension, loss of articulation devices, development of numerous nutritive foramina, and the tendency to fusion between the teeth in a tooth−family. Based on these shared characters, close phylogenetic relationships between the Protacrodontoidea, Hybodontoidea, and the Holocephali are postulated.
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The relatively rich assemblages of shark teeth from pelagic limestone (Mississippian, late Viséan, late Asbian-middle Brigantian) of three northern European regions: the Rhenish Mountains (Westenfeld Quarry, Germany), the Holy Cross Mountains (Todowa Grząba at the edge of Ostrówka Quarry, Poland), and Derbyshire (Cawdor Quarry, Matlock, England, UK) display certain similarities, with the absolute predominance of the teeth of Falcatidae (small Symmoriiformes) and the constant presence of Thrinacodus spp. The largest and most diverse assemblage from Todowa Grząba contains at least three species of a falcatid Denaea, a xenacanthimorph Bransonella nebraskensis, a newly described phoebodontid Thrinacodus dziki sp. nov., a few ctenacanthiform and euselachian teeth, and two abraded euchondrocephalan dental elements. Anachronistidae, common in the most of late Viséan pelagic faunas, are absent from Todowa Grząba and Westenfeld. The material under study differs from the shallow-water chondrichthyan fauna, hitherto described from the Mississippian carbonate platform facies, by its taxonomic content (particularly almost total absence of Euchondro-cephali), generally lower diversity, and higher frequency of small teeth.
The teeth of a well known late Palaeozoic cladodont chondrichthyan, “Cladodus” occidentalis from Russia, USA, and England are restudied and a new generic name, Glikmanius gen. nov., is proposed for this species. Yet another tooth−based species, formerly described as ?Symmorium myachkovensis, occurring on the Russian Platform and in Nebraska, is considered to belong to the newly erected genus. Although there is no direct evidence that Glikmanius possessed fin spines, the broad similarity between its teeth and those of Ctenacanthus compressus suggests it had a ctenacanthiform affinity. The possible relationships between Glikmanius, Cladodus sensu stricto, “Ctenacanthus” costellatus, and Heslerodus, are suggested. However, the proposition put forward by an earlier author that the teeth of Heslerodus might represent the lower jaw dentition of G. myachkovensis, is rejected. The overall resemblance of Glikmanius teeth and those of Cladoselache and Squatinactis is recognised as convergent.
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