The Ypresian Cambay Shale Formation of Vastan Lignite Mine in Gujarat, western India, has yielded a rich vertebrate fauna, including the earliest modern mammals of the Indian subcontinent. Here we describe its assemblage of four frogs, including two new genera and species, based on numerous, diverse and well−preserved ilia and vertebrae. An abundant frog, Eobarbourula delfinoi gen. and sp. nov., with a particular vertebral articulation similar to a zygosphene−zygantrum complex, represents the oldest record of the Bombinatoridae and might have been capable of displaying the Unken reflex. The large non−fossorial pelobatid Eopelobates, known from complete skeletons from the Eocene and Oligocene of Europe, is also identified at Vastan based on a single nearly complete ilium. An abundant “ranid” and a possible rhacophorid Indorana prasadi gen. and sp. nov. represent the earliest records of both families. The Vastan pelobatids and ranids confirm an early worldwide distribution of these families, and the bombinatorids and rhacophorids show possible origins of those clades on the Indian subcontinent.
The brown frogs (Rana temporaria-group) are a monophyletic group in the family Ranidae, which have a scarce fossil record in Europe that begins only in the Late Pliocene. A new fossil from the Dietrichsberg locality (Germany, Thuringia) extends their stratigraphic range back to the Early Miocene, and suggests that their origin lies outside the Western Palaearctic, most probably in Asia, with subsequent immigration in the Burdigalian.