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This paper describes the vertebrate ichnofauna from the Tumlin Sandstone (Buntsandstein) of the Holy Cross Mountains in Poland. The footprint assemblage has previously been regarded as Early Triassic in age; however, ichnogenera characteristic of the Late Permian are now recognized. Lack of representatives of the ichnofamily Chirotheriidae, characteristic of continental Triassic sediments worldwide, also indicates a Permian rather than a Triassic age for the studied assemblage. Three ichnogenera (Batrachichnus, Limnopus, and Amphisauropus) produced by amphibians are recognized, the remainder (Varanopus, Chelichnus, Dimetropus, Rhynchosauroides, Palmichnus, Paradoxichnium, and Phalangichnus) being of reptilian origin. Batrachichnus cf. salamandroides (Geinitz, 1861), Limnopus cf. zeilleri (Delage, 1912), Amphisauropus cf. latus Haubold, 1970, Varanopus aff. microdactylus (Pabst, 1896), Chelichnus cf. duncani (Owen, 1842), and Dimetropussp. are recorded in the Lower Buntsandstein for the first time. The following new ichnospecies are erected: Rhynchosauroides kuletae ichnosp. nov., Palmichnus lacertoides ichnosp. nov., Paradoxichnium tumlinense ichnosp. nov., Phalangichnus gradzinskii ichnosp. nov., and Phalangichnus gagoli ichnosp. nov.
We investigated the relation between the footprint shape of the fore and hind feet of sigmodontine rodents and their levels of arboreal activity. Footprint shape was obtained by analyzing the impressions left by identified animals captured in the field after being forced to pass through ink-tracking tunnels or by pressing their previously inked feet on a paper sheet. We used geometric morphometric techniques that use superimposition of landmarks (centers of the pads) to obtain footprint shape variables, which were reduced using multivariate analysis (principal component analysis). Arboreal activity was inferred on the basis of the proportions of individuals captured in arboreal traps (1.5–2.5 m height). Regression analysis of body size and the variable that best represented the footprint shapes (first principal component—PC1) did not indicate significant allometric effects on such shapes. We did not detect any significant phylogenetic effects on the arboreal activity of the rodents, either. The results indicated that the PC1 concerning footprint shapes of ten sigmodontine rodents efficiently reflects the degree of use of arboreal strata by these animals. The species studied showed different levels of arboreal activity and their hind footprints (r 2 = 0.94) were better indicators of arboreality than the fore footprints (r 2 = 0.53). These findings suggest a likely trade off for the fore feet functions. Such functions are probably not strictly related to locomotion. Other biomechanical functions (e.g., shock absorption) and/or manipulation (e.g., food manipulation and grooming) may exert relatively greater influence on the shape of fore feet.
In this study we analyzed daily pollen concentrations of Alnus spp. and Betula spp. from Worcester, UK and Wrocław, Poland. We analyzed seasonality, annual pollen index and footprint areas for the observed pollen concentrations by using the trajectory model hybrid single particle Lagrangian integrated trajectory (HYSPLIT). We examined 10 years of data during the period 2005–2014 and found substantial differences in the seasonality, pollen indices and footprint areas. For both genera, concentrations in Wrocław are in general much higher, the seasons are shorter and therefore more intense than in Worcester. The reasons appear to be related to the differences in overall climate between the two sites and more abundant sources in Poland than in England. The footprint areas suggest that the source of the pollen grains are mainly local trees but appear to be augmented by remote sources, in particular for Betula spp. but only to a small degree for Alnus spp. For Betula spp., both sites appear to get contributions from areas in Germany, the Netherlands and Belgium, while known Betula spp. rich regions in Russia, Belarus and Scandinavia had a very limited impact on the pollen concentrations in Worcester and Wrocław. Substantial and systematic variations in pollen indices are seen for Betula spp. in Wrocław with high values every second year while a similar pattern is not observed for Worcester. This pattern was not reproduced for Alnus spp.
In modern terrestrial ecosystems, the population size of large predators is low, and a similar pattern has usually been assumed for dinosaurs. However, fossil finds of monospecific, large theropod accumulations suggest that population dynamics were more complex. Here, we report two Early Cretaceous tracksites dominated by large theropod footprints, in Querulpa Chico (Peru) and Chacarilla (Chile). The two sites correspond to distinct depositional environments—tidal basin/delta (Querulpa Chico) and meandering river (Chacarilla)—with both subject to extensive arid or semiarid palaeoclimatic conditions. Although most trackways show no preferred orientation, a clear relationship between two trackmakers is observed in one instance. This observation, coupled with the high abundance of trackways belonging to distinct large theropods, and the exclusion of tracks of other animals, suggests some degree of grouping behaviour. The presence of freshwater sources in a dry climate and perhaps social behaviour such as pair bonding may have promoted interactions between large carnivores. Further, the occurrence of these two tracksites confirms that large theropod dinosaurs, possibly spinosaurids and/or carcharodontosaurids, existed on the western margin of Gondwana as early as the earliest Cretaceous.
Ichnotaxonomic revision of two extended sequences of large tetrapod footprints from the Westphalian A Bochum Formation of western Germany suggests assignment of the specimens to the well−known Permo−Carboniferous ichnogenera Ichniotherium and Dimetropus. Trackway parameters and imprint morphology strongly support basal diadectomorphs and “pelycosaurian”−grade synapsid reptiles, respectively, as the most likely trackmakers. The ichnofossils thereby extend the first appearance of these two important groups of basal tetrapods by about 5–10 million years, to the early Late Carboniferous, which is in accordance with the minimum age for the evolutionary origin of the clades following widely accepted phylogenetic analyses. These trackways provide not only direct evidence bearing on activity and behaviour of large terrestrial tetrapods close to the origin of amniotes, but also serve as a valuable benchmark for the assessment of controversially interpreted vertebrate tracks from other localities of similar age.
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