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Palaeontological events, documented by widespread beds or thin intervals of strata with either unusual (“exotic”) or acmes of common faunal elements are a characteristic feature of Upper Cretaceous epicontinental shelf sediments in NW Europe. Their importance in stratigraphic calibration has early been recognized and these “bioevents” are widely used as correlation tools. Furthermore, it appears that there is a genetic link between sequence and event stratigraphy as most of the “classic” bioevents developed during specific intervals of a 3rd−order depositional sequence. Early transgressive bioevents (ETBs) are subdivided into two subtypes, i.e., the lag and migration subtype. The lag subtype corresponds to the transgressive surface and develops in response to winnowing and relative enrichment of robust biogenic hardparts. Taphonomic alteration and time−averaging are important features. The migration subtype is related to the disappearance of physical or ecological barriers that triggered faunal migrations. Despite their onlapping character, most ETBs are quasi−isochronous, and their preservation potential is usually high. Thus, they are very useful stratigraphic markers. Maximum flooding bioevents (MFBs) represent autochthonous biogenic concentrations with relatively low shell densities. They are related to habitat stability and ecospace expansion, and develop by population blooms of taxa well adapted to the special maximum flooding conditions of the wide epicontinental shelf of NW Europe (e.g., low food availability). Cenomanian MFBs of NW Europe are not time−averaged and may comprise stratigraphically more expanded intervals with gradational lower and upper boundaries. Their often wide palaeogeographic extent associated with very high chances of preservation results in an excellent inter−basinal correlation potential. Late highstand bioevents (LHBs) are local to regional shell concentrations deposited as a result of increasing winnowing of fines and reworking by storms, currents and waves during late highstands. LHBs usually consist of paucior even monospecific skeletal concentrations with a high degree of fragmentation. Simple shell beds related to a single (storm) event, and composite (multiple−event) shell beds are recognized. LHBs share some features of ETBs, but lack of time−averaging, are laterally restricted and have low preservation potential. Thus, their importance in interbasinal correlation is poor. The time scales of Cenomanian bioevents range through several orders of magnitude (hours–days in LHB storm event concentrations to ~100 kyr in MFBs). In terms of position within sequences, the three bioevent types correspond to shell concentrations recognized in Mesozoic–Cenozoic formations around the world. Shell beds with similar positions within cycles as well as comparable sedimentologic and taphonomic characteristics have also been described from high−frequency sequences and parasequences, suggesting that the formational processes of shell beds operate in base−level controlled sedimentary cycles of different hierarchies (i.e., 3rd−up to 7th−order).
Gastropod response to a marked carbon isotopic geochemical anomaly across the Early–Middle Frasnian transition (Palmatolepis transitans–Palmatolepis punctata conodont zones) has been analysed along the southern Laurussian shelf, mainly within the Dyminy Reef in the Holy Cross Mountains. Gastropods are represented by three reefal associations (Kowalatrochus sanctacrucensis, Euryzone kielcensis, and Grabinopsis guerichi associations), and an impoverished open−shelf Straparollus laevis assemblage. The most severe diversity crisis is connected to the disappearance of local low−energy muddy habitats, as a result of a transgressive pulse (Middlesex Event) and benthic habitat changes tied to strongly fluctuating carbon cycling; this has been observed at the highly diverse Kadzielnia−type assemblage. Fifteen taxa have been recognised in this distinctive Early Frasnian mud−mound association, including six (probably endemics), which are unknown from the Middle Frasnian. The disappearance of three relict Givetian species (Euryzone delphinuloides, Straparollus laevis, and Goniasma? zarecznyi) is also recorded. Other species probably migrated into the shallower water part of Dyminy Reef and persisted in the Middle and Late Frasnian. The Middlesex Event and the earlier major biogeochemical perturbation seem to have less serious effects for evolution of gastropods in the Polish−Moravian part of the Laurussia shelf than the catastrophic Frasnian–Famennian extinction. Two new taxa are described: Frydiella kaimi gen. et sp. nov. (Eotomariidae) and Heidelbergeria czarnieckii gen. et sp. nov. (Elasmonematidae).
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