Paranyctoides aralensis, based on a partially known set of lower postcanines from the Turonian Bissekty Formation of Uzbekistan, is proposed to be a junior subjective synonym of Sailestes quadrans, itself based on an M1 from the same stratigraphic unit. As a result, the latter taxon is recombined as Paranyctoides quadrans comb. nov. Based on newly col− lected or recognized specimens, we show that this species has four lower incisors, including a procumbent i1 and a rela− tively small i4, as well as five double−rooted premolars, the smallest of which (p3) can be lost ontogenetically. The p5 of this species is premolariform, rather than molariform as previously thought based on an erroneous identification, and re− sembles North American species of Paranyctoides in the presence of a small, cingulid−like paraconid and a distal talonid cusp, as well as the absence of a metaconid. The lower molars of Paranyctoides are unique among Late Cretaceous eutherians in having a larger, lingually placed paraconid. Paranycoides probably is the sister taxon of the Late Cretaceous Zhelestidae.
Paranyctoides is represented by three named, and possibly four unnamed species in the Late Cretaceous, North America. P. aralensis from the Late Cretaceous of Dzharakuduk, Uzbekistan, belongs in this or a closely allied taxon. Lower molars have low trigonids, well-developed paraconids not appressed against metaconids, talonids on ml-2 as wide or wider than trigonids, hypoconulids often closer to entoconids than to hypoconids. Only two upper molars are known, both have comparatively narrow crowns with wide stylar shelves and stylar cusps, paracone and metacone separated, conules well developed, and protocone low. Pre- and postcingula vary from narrow in one, Sailestes quadrans, to wide in the other, Paranyctoides sp. Sailestes quadrans may be an metatherian. All known species of Paranyctoides from North America have a submolarifom ultimate premolar while Gallolestes pachymandibularis, also from North America, has molars not unlike those in Paranyctoides but may have an ultimate premolar with a molarifom trigonid. A specimen from Dzharakuduk referable to P. aralensis is suggestive of such morphology. At least P. aralensis had five premolars with the third reduced as in 'zhelestids'. These findings increase the Late Cretaceous North American/Asian ties even more for eutherians, now with 'zhelestids' and the Paranyctoides/Gallolestes clades known from both.
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