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Two new rhynchonelliformean brachiopods, Neumania paucicostata sp. nov. and Leoniorthis robusta gen. et sp. nov., are described from the Billingen and Volkhov stages (Lower–Middle Ordovician: Arenig) of the Baltic–Ladoga Klint area. All specimens were washed out from unconsolidated sediments, and were sampled in Tallin, Estonia, and at three localities in St. Petersburg district, Russia. Numerous specimens were found in a large mud mound, the lower part of which is within the Baltoniodus triangularis conodont Zone, otherwise usually missing in this area. L. robusta is here assigned to the family Orthidae.
A new species of the orthide brachiopod genus Kinnella is described from the Upper Member of the Georgian Bay Formation (Upper Ordovician) of Manitoulin Island, Ontario, Canada. This species, herein designated as Kinnella laurentiana sp. nov., occurs in strata of Richmondian (mid−Ashgill; Katian) age, most likely correlative with the eastern North American Dicellograptus complanatus Zone. This occurrence extends the known stratigraphic range of Kinnella downward considerably from its previously inferred basal Hirnantian inception. The new species is characterized by a moderately convex dorsal valve and an apsacline ventral interarea rarely approaching catacline. This is the third reported occurrence of Kinnella in North America, and is the only species known to have inhabited the epicontinental seas of Laurentia. The associated benthic shelly fauna indicates a depositional environment within fair weather wave base (BA 2). The ancestry of Kinnella and this species appears most likely to lie among older, morphologically similar members of the Draboviidae which were seemingly confined to higher latitude faunal provinces prior to the Hirnantian glacial event. Thus, the mid−Ashgill occurrence of Kinnella laurentiana in the palaeotropically located Manitoulin Island region suggests the mixing of a probable cooler water taxon with the warmer water epicontinental shelly fauna of Laurentia, as well as a possible earlier episode of low−latitude oceanic cooling. Cluster analysis of Kinnella−bearing brachiopod faunas reveals a sharp differentiation between the K. laurentiana−associated brachiopod fauna and all other known (Hirnantian–lower Rhuddanian) occurrences mainly represented by the type species K. kielanae.
This paper describes 37 species (4 new) belonging to 27 genera (1 new), 14 families, and 6 orders (Lingulida, Craniida, Strophomenida, Productida, Orthotetida, and Orthida) of silicified brachiopods from the middle Tournaisian (Mississippian, lower Carboniferous) of the vicinity of Gedongguan and Muhua villages (southern China). All specimens come from acid etching of detrital and oolitic limestone lenses scattered within grey to black laminated basinal micrite and marl of the Muhua Formation. The formation, which attains about 1–10 meters in thickness, is dated to the Siphonodella crenulata Zone. More than 10,000 silicified brachiopod specimens belonging to about 70 species were recovered from over 900 kg of the sampled limestone lenses, the most diverse brachiopod fauna of that age. The characteristic feature of the studied material is the prevailing disarticulation and fragmentation of skeletal parts due to their down slope transportation into a deeper water environment. Within Lingulida, one linguloid and one discinoid species are described. Craniida are represented by five species including Nematocrania pilea sp. nov. Strophomenida are represented by fragmentarily preserved specimens belonging to one species. The most diverse are Productida, which are represented by 7 chonetidine and 12 productidine species; new are Subglobosochonetes gedongguanensis sp. nov. and Globosochonetes gracilis sp. nov. New data on morphology of larval stage of Argentiproductus margaritaceus and its mode of attachment are presented. Orthotetida is represented by 5 species including Lamellispina spinosa gen. et sp. nov. Orthida is represented by 3, mostly cosmopolitan species. This study of a middle Tournaisian brachiopod fauna from Muhua, together with published data on the Mississippian brachiopods from other regions of South China, allow to study the Devonian–Carboniferous biotic crisis and post−crisis recovery.
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