The end of the carbonate sedimentation of the Famennian Wocklumeria limestone in the Holy cross Mts and Sudetes coincides with the disappemance of a high-diversity warm-water assemblage of ammonoids and conodonts with elaborated platform elements. In replacement, a low diversity ammonoid community of Acutimitoceras prorsum and a thin-crown conodont Protognathodus fauna migrated to the afea. When carbonate sedimentation was re-established in the Tournaisian, the new high-diversity ammonoid and conodont faunas represented again almost the whole range of morphologies known from the Famennian. Migrations into the area from unknown sources dominated, with little contribution from the local phyletic evolution. This characteristic ammonoid-conodont community disappeared with the sea-level rise in the Siphonodella crenulata Zone, to emerge at the same time in the North American Midcontinent. The reverse direction of migrations marks the latest Tournaisian Scaliognathus anchoralis event. In yet another cycle of the late Viséan, the new high-diversity faunas were not able to develop as elaborate conch or platform element morphologies as before. In a review of the literature it is shown how the Variscan orogenic activity, progressing towards the Northeast, and glaciations in Gondwana influenced the distribution of late Carboniferous ammonoids in Poland. Conodont taxa Weyerognathus gen. n., Neopolygnathus sudeticus sp. n., and Siphonodella belkai sp. n. are proposed.
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