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A new Late Carboniferous cnemidolestodean insect Xixia huban gen. et sp. nov., is described from the Xiaheyan locality (Zhongwei City, Ningxia, China; Tupo Formation). Its combination of character states and observed variants on the wing venations are indicative of homologies shared with more derived members of the order. In particular it is ascertained that MP runs fused with CuA and CuA + CuPa, but is not usually visible as a distinct vein. The new genus exhibits previously unknown coloration pattern composed of dark patches distributed over the whole forewing (as opposed to regular stripes or rows of spots, previously documented in the group). The comparatively abundant sample makes it one of the best documented cnemidolestodeans to date.
A new basal sphenacodontid synapsid, represented by an anterior portion of a mandible, demonstrates for the first time the presence of amniotes in the largest European Permo−Carboniferous basin, the Saar−Nahe Basin. The new taxon, Cryptovenator hirschbergeri gen. et sp. nov., is autapomorphic in the extreme shortness and robustness of the lower jaw, with moderate heterodonty, including the absence of a greatly reduced first tooth and only a slight caniniform development of the second and third teeth. Cryptovenatorshares with Dimetrodon, Sphenacodon, and Ctenospondylus, but notably not with Secodontosaurus, enlarged canines and a characteristic teardrop outline of the marginal teeth in lateral view, possession of a deep symphyseal region, and a strongly concave dorsal margin of the dentary. The new find shows that sphenacodontids were present in the Saar−Nahe Basin by the latest Carboniferous, predating the record of sphenacodontid tracks from slightly younger sediments in this region.
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Late Carboniferous bryozoans from La Hermida, Spain

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Fifteen bryozoan species belonging to thirteen genera have been identified from an outcrop of the Picos de Europa Formation (Moscovian, Upper Carboniferous) at La Hermida in northern Spain. Three species and one genus are new— Coscinium hermidensissp. nov., Cystodictya pustulosa sp. nov., and Cystocladia hispanica gen. et sp. nov. Rhabdomesid bryozoans are the most diverse order with seven species, followed by cystoporids (four species), fenestellids (three species) and trepostomids (one species). Bryozoans with erect branched or reticulate colonies dominate in the studied assemblage; only two species possess encrusting colonies. Together with associated crinoids, the bryozoan assemblage indicates a subtidal environment below the zone of vigorous water movement. The La Hermida bryozoan fauna confirms the Upper Carboniferous age of the Picos de Europa Formation and allows various biogeographical interpretations. All previously known species of the genus Coscinium were reported from the Lower Permian of Russia. Clausotrypa monticola is known from the Lower Permian of Russia and Arctic as well as from the Upper Carboniferous of Carnic Alps (Austria). Rhabdomeson cf. propatulissimum and Penniretepora pseudotrilineata are known from the same level of Italian Carnic Alps. Streblotrypa (Streblascopora) nikiforovae and Rhombocladia punctata are known from the Upper Carboniferous (Moscovian) of Ukraine. Fistulipora petaloida is known from Kasimovian Stage of Russian Plate. Several other species show connections with North America.
Ichnotaxonomic revision of two extended sequences of large tetrapod footprints from the Westphalian A Bochum Formation of western Germany suggests assignment of the specimens to the well−known Permo−Carboniferous ichnogenera Ichniotherium and Dimetropus. Trackway parameters and imprint morphology strongly support basal diadectomorphs and “pelycosaurian”−grade synapsid reptiles, respectively, as the most likely trackmakers. The ichnofossils thereby extend the first appearance of these two important groups of basal tetrapods by about 5–10 million years, to the early Late Carboniferous, which is in accordance with the minimum age for the evolutionary origin of the clades following widely accepted phylogenetic analyses. These trackways provide not only direct evidence bearing on activity and behaviour of large terrestrial tetrapods close to the origin of amniotes, but also serve as a valuable benchmark for the assessment of controversially interpreted vertebrate tracks from other localities of similar age.
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