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The catchment of the River Kokemäenjoki covers ca. 27 100 square kilometers in western Finland, and the the length of the river is ca 120 km. The river discharges into the Bothnian Bay, the northern section of the Baltic Sea. The delta is changing and prograding towards the sea exceptionally rapidly. The pace of the growth of the deltaic formations,a s well as the major zones of the macrophytivc vegetation is nowadays some 30–40 meters a year. This makes the delta the most rapidly changing aquatic and littoral ecosystem in the Northern Europe. The Kokemäenjoki River delta is often characterized as a biological hotspot, and major sections of the estuary are included in several leading international nature conservation programmes, i.e. The Natura 2000 network of the European Union, the intergovernmental The Ramsar Convention on Wetlands, as well as in the IBA, Important Bird Area programme established by the BildLife International organization. The diversities of both flora and fauna are very high – at least as regards of the northern location of the estuary. In the Kokemäenjoki River ca. 420 taxa of flowering plants (Spermatophyta) are permanently found (i.e. about a third of the ca. 1 200 flowering plants permanently growing in Finland). Of the pteridophytes (Pteridophyta), there are 17 out of the 57 species of the Finnish taxa growing at the delta. Of the stoneworts (Chrarophyta) the delta is home of 9 species of the 21 taxa found in Finland. The number of bird species breeding in the area is ca. 110, and the total number of bird taxa living and resting during the migration periods amounts to 220 species (i.e. nearly the whole range of the 248 bird species of the permanent Finnish bird taxa). Of the invertebrates, the species composition of dragonflies (Odonata) is best known. There are 25 species of the 55 dragonfly taxa found in Finland. Of the plant species occupying the delta, 5 species of flowering plants and one species of stoneworts are classified as endangered, fulfilling the red list criteria established by the International Union for Conservation of Nature (IUCN). Of the bird species of the estuary, 31 species are included in the Red List of endangered species. The present paper summarizes the data presented in various reports, mainly in Finnish, and majority of them in hardly accessible depositories.
The growth dynamics of two tall littoral helophytic plants, the narrow-leaved cattail (Typha angustifolia L.) and broad-leaved cattail (Typha latifolia L.; Typhaceae) were studied in the rapidly changing estuarine habitats in the Kokemäenjoki River delta, western Finland. The two cattails form uniform, single-species communities (monocultures) throughout the plant-covered estuary. Of the two taxa compared, the shoots were taller in T. angustifolia (mean 166 cm) than in T. latifolia (mean 120 cm). But due to the robust leaves, the relation in the average weight of individual ramets was opposite: The mean weight of T. angustifolia was 9.6 g (dry wt), and that of T. latifolia was 16.5 g. In a separate study, the leaf height was compared between the fertile (flowering) and sterile (non-flowering) ramets. In flowering ramets the average leaf length was 35 cm taller in Typha angustifolia than in T. latifolia. The differences were even more pronounced in sterile ramets, where the leaves of Typha angustifolia were 70 cm taller than those of T. latifolia. The differences were statistically highly significant. Interspecific competition between the two Typha species is negligible, because the microhabitats differ from each other. T. angustifolia grows in considerably deeper (mean depth 42 cm) waters than T. latifolia (mean depth 19 cm). The optimum range in the water depth is markedly stricter in T. angustifolia than in T. latifolia. The differences between the rooting depths of the two cattails were statistically highly significant. The physico-chemical characteristics of the rooting zones (rhizospheres) of the two cattails are similar, with the locally produced (autochthonous) organic matter dominating and determining the fertility of the habitats.
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