Bicentric versus monocentric concept of Tabaninae evolution are discussed. Recent division of the subfamily, including redefinition of the tribe Lepidoselagini, opened the possibility of tracing plesiomorphic taxa In Northern Hemisphere. 9 such genera with 26 species are distributed in Holarctic. They occur chiefly in arboreal centers. The disjunction of Glaucops Szil. and partly of Ochrops Szil. indicates preglacial occurrence of the tribe in Northern Hemisphere.