Development of microspores and pollen grains of Cucumis sativus L. was studied at the light microscope level from meiosis to mature pollen. The first pollen mitosis occurs after microspore release from the tetrad at the vacuolate microspore stage, but this stage of development is not correlated with enlargement of the entire pollen grain; pollen grain volume grows continuously from microspore release until pollen dehydration, simultaneously with an increase in the amount of cytoplasm in the vegetative cell. Starch grains in the vegetative cell appear after displacement of the generative cell inside the pollen grain and disappear in the final stage of pollen maturation. Mature pollen grains are bicellular at anthesis.