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Five Lower Carboniferous species of Cyrtosymbolinae Hupé, 1953 are redescribed and one new is established. They come from the territory of Eurasia and represent two rare cyrtosymbolinid genera: Weania Campbell, 1963 and Griffithidella Hessler, 1965; one of them is tentatively assigned to the genus Phillibole Richter & Richter, 1937.
Snout is very deep and laterally compressed in the Protoceratopsidae. Oral cavity is highly vaulted longitudinally and transversely. It wedges into the middle of the nasal chamber dorsally causing that ventral portion of the latter parallels oral cavity on both its sides. Fenestra exochoanalis is oriented almost vertically, opening on the side of the oral cavity. Structure of the snout and hard palate results in narrowing and deepening of the narial passage, thus inhaled air passed close to large surface of the mucosa — covered walls of the nasal cavity. Blood cooling effect of that narrow air passage may have been significant in the Protoceratopsidae. Choana opened well in front of the entrance to larynx: separation of feeding from breathing was not complete, although extensive.
The new Famenrtian triolobite species Trimerocephalus dianopsoides including the meraspid period is described. The mode of moulting in young and adult phacopids and phylogenetic relations of the genera Cryphops R. & E. Richter, Trimerocephalus M'Coy and Dianops R. & E. Richter are discussed.
Famennian and Lower Carboniferous trilobites of the subfamily Cyrtosymbolinae (Proetidae) from the cephalopod biofacIes of the Holy Cross Mountains (Góry Świętokrzyskie) are described. Of the 51 species described, 20 are new and 26 are identified only at the generic level. The variability and ontogenetic development of the species are investigated. Suggestions on the occurrence of proetid and phacopid trilobites in correlation with the type of lithofacies are made. An attempt is made to base the stratigraphy of Famennian and Lower Carboniferous beds on the trilobites here studied.
An incomplete skeleton of a theropod dinosaur, Bagaraatan ostromi gen, et sp. n., was found in the Nemegt Fm. at Nemegt, Mongolia. The mandible in B. ostromi has a shallow but massive dentary, relatively deep postdentary portion with two surangular foramina and somewhat elongated retroarticular process; on the lateral surface of the postacetabular process of the ilium there are two large depressions for muscle origins separated by a crestlike projection; the fibula is fused distally with the tibiotarsus and the coalesced astragalocalcaneum. Bagaraatan represents the Tetanurae and displays some synapomorphies with the Avetheropoda, however, incompleteness of the skeleton of B. ostromi does not allow to determine its more precise affiliation. Bagaraatan was about 3.0-3.5 m long, had a relatively small head and slender hind limbs. The presence of strongly developed hyposphenes in a long series of anterior caudals rendered its tail only slighily flexible proximally.
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Nasal salt gland in dinosaurs

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Large, herbivorous dinosaurs display, as a rule, enlarged external nostrils. It is considered here, that they accomodated large functioning salt gland homologous with the lateral nasal gland of Recent reptiles and birds. All dinosaurs were probably uricotelic animals which had to use the extrarenal way of excreting excess of monovalent ions. It is suggested here that they were able to use the nasal salt gland for this purpose. Its presence may have been especially important for unloading the excess of potassium ions ingested by large herbivores with their vegetarian food, or/and for getting rid of sodium ions - by herbivores living in saline environment. An alternative is also given, that the development of large functioning salt gland may have been exclusively a result of large body size and consequently of large amount of potassium ions ingested, independent of fresh water availability in the environment.
Three Lower Carboniferous representatives of the family Otarionidae: Coignouina acanthina (Coignou, 1890), Namuropyge discors discors (M'Coy, 1844), N. discors kingi R. & E. Richter, 1939 are here redescribed and figured. Their systematic position and the constitution of the family Otarionidae are discussed. Reconstructions of Coignouina acanthina and Namuropyge discors discors are given.
This paper deals with 13 species of Couvinian Trilobites from Wydryszów. In addition to species previously recorded from the Holy Cross Mts, the writer describes some others not known so far from Poland and 3 new species.
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Nine species of the Famennian phacopid trilobites are described from the southern part of the Holy Cross Mts. In addition to forms already recorded from this area the writer describes some species never before found in Poland, as well as two quite new species, namely: Trimerocephalus polonicus and Dianops? trifolius.
Two new trilobites: Ditomopyge roemeri spitsbergensis n. subsp. and Paladin trigonopyge n. sp. are described. D. roemeri spitsbergensis seems to indicate the Upper Carboniferous age for these beds, as its close relatives D. roemeri roemeri (v. Moeller, 1867) and D. grünewaldti (v. Moeller, 1867) are known from the Gzhelian Stage of the Urals, Donetz Basin and Voronezh Region (USSR).
Seven species and subspecies of the Lover Carboniferous Cyrtosymbolinae Hupé from Poland and Great Britain are described, including 5 new ones. They are assigned to 5 subgenera (one new) of the genus Archegonus Burmeister, 1843 (emend. Hahn, W65). Archegonus (Cyrtoproetus) Reed, 1943 is revised. The relation between Archegonus (Phillibole) R. & E. Richter, 1937 and Liobole R. & E. Richter, 1949 is discussed. Also described is the ontogenetic development of Archegonus (Phillibole) aprathensis richteri n.subsp.
Several Carboniferous representatives of Brachymetopus McCoy are described from Poland and U.S.S.R. They are: B. maccoyi maccoyi (Portlock, 1843), B. weberi n. sp., B. ouralicus sanctacrucensis n. subsp., B. moelleri parvus n. subsp., Brachymetopus n. sp. and ?Brachymetopus sp. The widely distributed species B. ouralicus (de Verneuil, 1845) is revised.
A new theropod dinosaur Borogovia gracilicrus gen. et sp. n. assigned to Troodontidae is described, based on fragmentary hind limbs of one individual from the Upper Cretaceous Nemegt Formation of Gobi Desert, Mongolia. A unique feature of B. gracilicrus is a straight ungual in the second, specialized toe of the pes. The known skeletal elements of all troodontid species are listed (Table 1). Taxonomic status of Saurornithoides junior is considered.
Brains in living tetrapods other than birds and mammals do not entirely fill the brain cavities. Examination of dinosaur braincases does not usually allow determination relating to how close walls of endocranial cavity lay to the surface of brain. The here described fragment of a skull roof of an oviraptorid dinosaur, Ingenia yanshini, shows perfectly preserved, numerous vascular imprints that cover the internal surfaces of frontals and parietals in the region roofing the cerebral hemispheres and cerebellum. This specimen shows that in oviraptorids the brain closely fitted the brain cavity, to the extent found in birds and mammals. Among dinosaurs, only one similar case has been previously reported in an ornithomimid, Dromiceiomimus brevitertius, but the preserved vascular imprints are less numerous and regular in this dinosaur than in Ingenia yanshini.
The well preserved material of the Late Cretaceous dromaeosaurid, Velociraptor mongoliensis, has allowed us to supplement earlier descriptions of the skull in this species. The skull of V. mongoliensis is similar to that of Deinonychus antirrhopus, but differs from the latter by: (1) laterally convex supratemporal arcade resulting in short, rounded supratemporal fenestra; (2) depressed nasal; (3) longer maxillary process of premaxilla; (4) lack of separate prefrontal, and (5) convex ventral border of the dentary. These differences, especially that in the structure of the temporal region, support generic distinction of Deinonychus and Velociraptor. Skulls of other dromaeosaurids are compared.
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On ornithischian phylogeny

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The ornithischian dinosaurs are analysed using the cladistic methodology. The preferred hypothesis (see fig. 1) is that: 1. Ornithischia are monophyletic, 2. Ankylosauria are sister group of all other ornithischians, 3. Pachycephalosauria + Ceratopsia are monophyletic unit sharing common characters not found in other ornithischians. Quadrupedality is considered as primitive ornithischian condition.
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The quadrate of oviraptorid dinosaurs

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The quadrate of oviraptorid dinosaurs is strongly pneumatized and differs from the quadrates of other non-avian theropods by: (1) two separate facets on the otic process for contacts with the squamosal and braincase; (2) the articular surface for the pterygoid extended to the articular surface of the medial mandibular condyle; (3) the mandibular process provided laterally with a quadratojugal process bearing the quadratojugal cotyla. In the above characters the oviraptorid quadrate resembles those in most ornithothoracine birds, but, contrary to the streptostylic quadrate of birds, the oviraptorid quadrate is monimostylic.
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All articulated hadrosaurian skeletons expose a comparatively abrupt ventral flexure of anterior thoracal portion of the vertebral column. This resulted in effective shortening of the body anterior to the acetabulum and in shifting backward the gravity center of the heavy thoracal part, in deepening of pleural cavity as well as in lowering the suspension point of the fore limb at the glenoid, making easier access of the fore limb to the ground. In some hadrosaurs at least, sacropelvic contact was strengthened by a forward extension of the acetabular bar, so that pubis also was keyed to sacrum by a sacral rib, and the puboiliac contact was reinforced. This enabled these hadrosaurs to assume and sustain a stance with the vertebral column inclined upwards.
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