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A pattern of wolfCanis lupus Linnaeus, 1758 predation on red deerCervus elaphus Linnaeus, 1758 was studied in Bieszczady Mountains in 1991–2002. In total 324 remains of red deer > 4 months old, killed by wolves throughout the year, were found. The sex, age and bone marrow fat content of wolf kills were compared with the same characteristics within the free living red deer population. The overall contribution of calves killed by wolves (24%) in October-May was higher than in the population (17%), and decreased from autumn to spring. Adult males were more vulnerable to wolf predation than adult females: stags constituted 62% and hinds 38% of adult red deer killed by wolves, whereas in the population, the percentages were 37 and 63%, respectively. Stags killed by wolves were younger (ˉx=4.1 years old) than hinds (ˉx=8.9 years old). Wolves killed more > 8 years old hinds and < 5 years old stags than available in the population. In wolf kills, the average fat content in femur marrow was higher among hinds (84.9%) than stags (69.3%) and calves (66.1%). Only 8% of hinds had < 70% femur marrow fat content, whereas 40% of calves and 38% of stags had marrow fat values below that level. Marrow fat content showed seasonal variation and was the lowest in March among all sex-age classes. The monthly share of stags in all kills, and hinds in hind-calf part of the sample was negatively correlated with their monthly average bone marrow fat content, and monthly share of calves was positively correlated with monthly average bone marrow fat content of adults. The segregation of social units (hind-calf and stag groups), except during the rutting season, and the low fat reserves of males from midwinter until spring contribute to the high overall incidence of calves and adult males and the relatively low incidence of adult females among wolf kills.
From 1991 to 1995, wolf Canis lupus (Linnaeus, 1758} population dynamics were studied in Bieszczady National Park and the surrounding area (520 km2). The study area was utilized by 5 packs, Pack sizes averaged 5.6 in early and 3.9 in late winter. Overwinter declines in wolf numbers ranged from 21% to 39% (x = 29%), which corresponded well to the known number of wolves killed by hunters or dead of other causes. After every winter decline, wolf numbers recovered through reproduction. Generally, wolf numbers were stable or slightly decreasing during the study. Three neighbouring wojf packs occupied an area of 340 km" and the estimated territory size averaged 85 km2. The estimated density of wolves averaged 5,1/100 km2 in early winter and 3.3/100 km' in late winter. Of all known causes of wolf mortality, 86% were from legal hunting, 5% were from poaching, and 9% were from natural causes. Bieszczady National Park is small in size and its topography influences the spatial distribution of packs. No single pack was fully contained within, or protected by the Park. The number of wolves is overestimated in official reports, because the same packs are likely counted as different groups in neighbouring census units. On hunting grounds adjacent to Bieszczady NP, harvest plans exceed the actual number of wolves which inhabit the area. The creation of a wolf protection zone around Bieszczady NP and some regulations for wolf management in the rest of the region are proposed.
The diet of wolves Canis lupus Linnaeus, 1758 was studied from October 1989 to November 1992 in the Bieszczady Mountains, southeastern Poland. A total of 221 wolf scats were collected and analyzed to determine the prey species consumed by wolves in each season. Cervids (red and roe deer) obviously predominated in wolf diet and consisted from 65% of winter biomass to 96% of summer biomass consumed. The red deer made up approximately 95% and roe deer only 5% of total cervid biomass con­sumed. During summer deer fawns made up 28% of total cervid biomass consumed. The wild boar constituted more significant food only during winter - 17% of biomass eaten. Among wild boars, piglets were selected and made up 66% of total wild boar biomass consumed during winter. Domestic livestock represented more significant food in winter (16% of biomass eaten) and was consumed as carrion laid out by hunters to bait wolves. Other food categories as hares, voles and insectivores played a negli­gible part in the wolf diet. Spring, summer and autumn diet were little diversified. Only winter diet differed significantly from other seasons for the presence of wild boar and cervids.
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