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A tiny tarsioid primate occurs in early Eocene sediments of the Naran Bulak Formation, southern Gobi Desert, Mongolian People's Republic. The new primate, Altanius orlovi, new genus and species, is an anaptomorphine omomyid and therefore belongs to a primarily American group of primates. Altanius is apparently not a direct ancestor of the Asian genus Tarsius. American rather than European zoogeographic affinities are indicated, and this in turn supports the view that for a time in the earliest Eocene the climate of the Bering Route was sufficiently warm to support a primate smaller than Microcebus.
We describe Dissacus zanabazari new species from a partial skeleton collected from the early Eocene Bumban Member of the Naran Bulak Formation at Tsagaan Khushuu (Omnogov Province, Mongolia). The holotype includes most of the skull with basicranium, mandibles, well preserved upper and lower dentitions, partially articulated left manus and right tarsus, and most of the long bones in the limbs. The presence of a rudimentary 1st metatarsal is confirmed in mesonychids, and the relatively unworn lower incisors display an unexpected trilobed morphology. Autapomorphies of this new species are short face, absence of diastemata between the lower premolars (except between p1 and p2), m3 metaconid subequal to protoconid, and foramen for superior ramus of stapedial artery entirely within the petrosal. A phylogenetic analysis of 89 characters scored for 14 mesonychians and 5 outgroups resulted in 8 most parsimonious trees. Dissacus zanabazari is in a clade with D. navajovius, but this genus is otherwise paraphyletic. The strict consensus of the eight trees has a monophyletic Mesonychia, Hapalodectidae, and Mesonychidae; Dissacus and Ankalagon as the most basal mesonychid genera; and paraphyly of Pachyaena.
We describe a partially crushed skull and dentaries of a sub-adult individual of Daulestes nessovi sp. n., from the Coniacian of Uzbekistan. This is the earliest known eutherian skull (about 87 Ma) and the sixth genus of Cretaceous eutherians in which a skull is available. Because the skull of D. nessovi is sub-adult, certain plesiomorphic features may be ontogenetic and should be interpreted with caution. Four upper premolars and five lower premolariform teeth were in use (possibly to become four lowers when fully adult). The upper cheek-teeth have winged conules; M2 has large parastylar and small metastylar projections. Pre- and postcingula are lacking on DP4 and the upper molars. The talonids of dp4-m2 are about 90% as wide as the trigonids, with widely separated entoconid and hypoconulid. The skull has a large sphenorbital fissure, no foramen rotundum, and apparently no pterygoid process of the sphenoid. A large orbital wing of the palatine prevents maxilla-frontal contact within the orbit. The zygomatic arch is slender. The cochlea has one full turn, with an expanded apex, which suggests that a lagena might have been present. A large malleus with a robust anterior process, and a large promontorium may be due to young age of the individual or a primitive retention, as in the platypus. Because of the similarity to Asioryctidaem both cranial structure and dentition, we assign Daulestes tentatively to Asioryctitheria Novacek et al. 1997, family incertae sedis.
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