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This paper is the second in a series designed to cover taxonomically all Australian species of Diomus Mulsant. Nomenclatural history, diagnoses, digital illustrations and distribution maps are provided for each species. Thirty species are treated in this paper, among those 14 are new: D. bunya sp. nov., D. carbine sp. nov., D. circus sp. nov., D. gingera sp. nov., D. gilvus sp. nov., D. hebes sp. nov., D. kioloa sp. nov., D. leai sp. nov., D. lord sp. nov., D. micrus sp. nov., D. pisinus sp. nov., D. prodigialis sp. nov., D. tasmanicus sp. nov. and D. villus sp. nov. Diomus hypocrtus Weise, 1923 is synonymised with Scymnus mareebensis Blackburn, 1895 syn. nov. Lectotypes are designated for the following taxa to stabilize their taxonomic positions: Diomus hypocrtus Weise, 1923; Scymnus cowleyi Blackburn, 1895; S. elutus Lea, 1902; S. flavifrons var. norfolcensis Lea, 1929; S. frater Lea, 1902; S. impictus Blackburn, 1895; S. inaffectatus Blackburn, 1892; S. insidiosus Blackburn, 1889; S. macrops Lea, 1929; S. maestus Lea, 1926; S. mareebensis Blackburn, 1895; S. obumbratus Blackburn, 1895; S. sublatus Blackburn, 1892; S. triangularis Lea, 1902; S. whittonensis Blackburn, 1892; S. victoriensis Blackburn, 1892 and S. yarrensis Blackburn, 1895.
The first paper dealing with revision of the Australian members of cosmopolitan genus Diomus Mulsant treats 30 species, mostly with distinct colour pattern, that can be identified using external characters. Nomenclatural history, diagnoses, illustrations and distribution are provided for each species. Fifteen new species are described: Diomus ancorus sp. nov., D. bimaculatus sp. nov., D. brookfieldi sp. nov., D. capital sp. nov., D. kuranda sp. nov., D. kosciuszko sp. nov., D. millaamillaa sp. nov., D. marmorosus sp. nov., D. ningning sp. nov., D. reidi sp. nov., D. storeyi sp. nov., D. tinaroo sp. nov., D. torres sp. nov.,D. weiri sp. nov., D. zborowskii sp. nov.. The following new synonyms are proposed: Diomus sphragitis (Weise, 1885) = Scymnus trilobus Lea, 1902 = Scymnus indistinctus Lea, 1902; Diomus Sydneyensis (Blackburn, 1892) = Scymnus mimicus Lea, 1902 = Scymnus pectoralis Lea, 1902 (new synonyms). Scymnus australis Blackburn, 1889 is transferred to Diomus (new combination). Lectotypes are designated for the following taxa: Diomus rusticus Weise, 1859; Scymnus australis Blackburn, 1889; S. corticalis Lea, 1908; S. cucullifer Blackburn, 1892; S. ementitor Blackburn, 1895; S. flavifrons Blackburn, 1889; S. flavolaterus Lea, 1926; S. indistinctus Lea, 1902; S. jocosus Blackburn, 1892; S. kamerungensis Blackburn, 1895; S. meyricki Blackburn, 1889; S. mimicus Lea, 1902; S. notescens Blackburn, 1889; S. pectoralis Lea, 1902; S. (D.) pumilio Weise, 1885; S. (D.) scapularis Weise, 1885; S. striatus Lea, 1902; S. sydneyensis Blackburn, 1892; S. tenebricosus Boheman, 1859 and S. trilobus Lea, 1902.
The Oriental genus Macroilleis Miyatake is revised. Three species are recognized, including one new species Macroilleis borneensis sp. nov. (Borneo, Sabah). All species are redescribed and illustrated. A key to the species is given.
We studied molecular and morphological diversity of Platerodrilini from China. Oriental and Palearctic Platerodrilus were included in a phylogenetic analysis and major lineages are identified using molecular data and morphology. We reinstate Duliticola Mjöberg, 1925 from the synonymy of Platerodrilus Pic, 1921 as a subgenus in Platerodrilus s. 1. Additionally, we propose a new subgenus Zhelongia subgen. nov. for the type-species Platerodrilus (Zhelongia) igneus sp. nov. and three related species. Larva of P. (Z.) igneus sp. nov. is described. Zhelongia subgen. nov. represents a deeply rooted lineage in Platerodrilus s. 1. and all species from China, India, and Indo-Burma are included in Zhelongia. The highest diversity of Platerodrilus is found in the Sundaland and Chinese species represent a species-poor but morphologically distant lineage. The ranges of Chinese species define the northernmost distribution limits of Platerodrilus. These limits are probably affected by climatic fluctuations and the low ability of neotenics to rapidly colonize distant areas during interglacial periods. Lizhongiella gen. nov., a new genus of Platerodrilini is described for Lizhongiella hainanensis sp. nov. from Hainan.
The Australian species classified in the cosmopolitan genus Scymnus are revised. Nomenclatural history, diagnoses, illustrations and distribution are provided for each of the eleven recognized species. Scymnus fuscatus Boheman, 1859 is recorded from Australia for the first time. Four new species, Scymnus alligator, S. bunya, S. leai and S. tasmanicus are described. Lectotypes are designated for: Midus pygmaeus Blackburn, 1892, Scymnomorpha duplopunctulata Blackburn, 1892, Scymnus ambulans Blackburn, 1895, S. mitior Blackburn, 1895, S. parallelus Blackburn, 1889, S. queenslandicus Blackburn, 1892 and S. varipes Blackburn, 1895. A new name Scymnus blackburni nom. nov. is introduced for Scymnus pygmaeus (Blackburn, 1892) preoccupied by Scymnus pygmaeus (Fourcroy, 1785). Scymnus varipes Blackburn, 1895 is recognised as a junior synonym of Scymnus ambulans Blackburn, 1895 (syn. nov.)
The tribe Lyponiini is studied with morphology and seven new species from China are described: Lyponia (Lyponia) cangshanica sp. nov., L. (L.) hainanensis sp. nov., L. (Poniella) pulchella sp. nov., L. (P.) qinlingensis sp. nov., L. (P.) tianquanensis sp. nov., L. (Weiyangia) muyuensis sp. nov. and Ponyalis variabilis sp. nov. The diagnostic characters are described and illustrated.
The Australian Dicranolaius fauna is revised based on 79 described species (30 of them are new). The revision is mainly based on males as they only provide good diagnostic characters for species recognition. The following new species are described: Dicranolaius acacia sp. nov., D. alice sp. nov., D. anic sp. nov., D. archer sp. nov., D. balah sp. nov., D. bogan sp. nov., D. brittoni sp. nov., D. cardaleae sp. nov., D. chinchilla sp. nov., D. confusus sp. nov., D. crypticus sp. nov., D. desertus sp. nov., D. eromanga sp. nov., D. hudsoni sp. nov., D. kildurk sp. nov., D. kimberley sp. nov., D. kinchega sp. nov., D. micrus sp. nov., D. moffatt sp. nov., D. monteithi sp. nov., D. pilbara sp. nov., D. proserpine sp. nov., D. pulleni sp. nov., D. quorn sp. nov., D. reidi sp. nov., D. rockhampton sp. nov., D. spinifex sp. nov., D. thylungra sp. nov., D. uptoni sp. nov., D. zborowskii sp. nov. The following new combinations are proposed: D. ammophilus (Lea, 1917) (Laius), comb, nov., D. distortus (Blackburn, 1888) (Laius), comb, nov., D. egenus (Lea, 1899) (Laius), comb, nov., D. eremita (Blackburn, 1895) (Laius), comb, nov., D. eyrensis (Blackburn, 1892) {Laius), comb, nov., D. guttulatus (Fairmaire, 1877) (Laius), comb, nov., D. melanoderes (Lea, 1917) (Laius), comb, nov., D. minutus (Lea, 1914) (Laius), comb. no\., D.purpureiceps (Lea, 1915) (Laius), comb. nov.,Z>. variegatus (Blackburn, 1889) (Laius), comb, nov., D. verticalis (Macleay, 1826) (Laius), comb, nov., Intybia cavicomis (Lea, 1909) (Laius), comb, nov., Intybia hackeri (Lea) (Laius) comb, nov.; Simoderus megalops (Lea) (Laius) comb. nov. The following synonymies are proposed: D. bellulus (Boisduval, 1835) [= D. cyanocephalus (Lea, 1909); = D. intermedius (Lea, 1909); =D. nidicola (Lea, 1909); =D. orcicornis (Lea, 1909)],D. conicicornis (Blackburn, 1888) [= D. rugiceps (Lea, 1899)], D. guttulatus (Fairmaire, 1877) [= D. longus (Lea, 1929)], D. verticalis (Macleay, 1826) [ = D. tarsalis (Lea, 1909)]. The lectotypes of the following species are designated: Laius acervatus Lea, 1917; L. ammophilus Lea, 1917; L. aulacophoroides Lea, 1917; L. c-pupureus Lea, 1914; L. concavifrons Lea, 1917; L. conicicornis Blackburn, 1888; L. curvicornis Lea, 1929; L. cyanocephalus Lea, 1909; L. distortus Blackburn, 1888; L. egenus Lea, 1899; L. eremita Blackburn, 1851; L. eyrensis Blackburn, 1892; L. fimbriceps Lea, 1929; L. flavifrons Lea, 1917; L. flavonotatus Lea, 1917; L. guttulatus Fairmaire, 1877; L. inconstans Lea, 1921; L. intermedius Lea, 1909; L. intricatus Lea, 1921; L. longus Lea, 1929; L. major Blackburn, 1889; L. mastersii Macleay, 1872; L. melanoderes Lea, 1917; L. minutus Lea, 1914; L. nidicola Lea, 1909; L. orcicornis Lea, 1909; L. orthodoxus Lea, 1909; L. pallidus Lea, 1899; L. plagiaticollis Fairmaire, 1877; L. planiceps Lea, 1899; L. purpureiceps Lea, 1915; L. rugiceps Lea, 1899; L. sinus Lea, 1899; L. tarsalis Lea, 1909; L. tetrasticus Lea, 1922; L. trifoveicornis Lea, 1921; L. variegatus Blackburn, 1889; L. villosus Lea, 1899; Malachius verticalis Macleay, 1826. Neotype oiMalachius bellulus Boisduval, 1835 is also designated.
Phylogenetic relationships within the family Rhipiceridae were investigated to elucidate the relationships between genera and subgenera of Rhipicerinae. Eleven ingroup taxa and three outgroups were included in cladistic analysis, based on 34 characters derived from adult morphology. Phylogenetic analysis confirms that Rhipicerinae are monophyletic and may be divided into four genera: Oligorhipis (stat. nov.), Polymeriiis, Polytomus (stat. rev.) and Rhipicera. All genera of Rhipicerinae are thoroughly described, illustrated and key to their identification is provided. Species level reviews and keys are provided for Oligorhipis and Rhipicera. Two new species are described: Oligorhipis vanemdeni sp. nov. (Western Australia) and Rhipicera carinata sp. nov. (Western Australia). Rhipicera abdominalis Klug, 1825, R. cyanea Castelnau, 1834, R. dalmanni Westwood, 1837 and R. marginata Kirby, 1818 are transferred to Polytomus Dalman (comb. nov.). Agathorhipis bifossata Fauvel, 1904 and Ptiocerus vestitus Castelnau, 1840 are transferred to Oligorhipis Guérin-Méneville, 1843 (comb, nov.). Agathorhipis Guérin-Méneville, 1843 is synonymised with Rhipicera Latreille, 1817 (syn. nov.) and Rhipicera (Agathorhipis) neglecta Emden, 1925 with Rhipicera femorata Kirby, 1819 (syn. nov.). The neotype of Rhipicera reichei Guérin-Méneville, 1843 is designated to stabilize the taxonomie identity of this species.
A new genus, Cretodascillus gen. nov. in the family Dascillidae with a new species C. sinensis sp. nov., is described from the Early Cretaceous Yixian Formation of Inner Mongolia, China. The fossil taxa described in Dascillidae have been critically reviewed and the Upper Triassic Apheloodes Dunstan, 1923 (type species A. obliquum Dunstan, 1923) and Leioodes Dunstan, 1923 (type species L. planum Dunstan, 1923) are removed from Dascillidae to Coleoptera incertae sedis, the Miocene Protoacnaeus Wickham, 1914 (type species P. tenuicomis Wickham, 1914) is transferred to Psephenidae and Miocyphon Wickham, 1914 (type species M. punctulatus Wickham, 1914) to Scirtidae.
Phylogenetic relationships within the Dascillinae were investigated. Eighteen ingroup taxa and two outgroups were included in cladistic analysis, based on 30 characters derived from adult morphology. Six genera are recognised in Dascillinae: Coptocera Murray, Dascillus Latreille, Metallidascillus Pic, Notodascillus Carter, Petalon Schoenherr and Sinocaulus Farimaire. Pseudolichas Fairmaire, 1878 (TS: P. sulcifrons Fairmaire, 1878) is recognised as a junior synonym of Petalon Schoenherr, 1833 (TS: Bruchus fulvulus Wiedemann, 1819) syn. nov. All valid genera of Dascillinae are thoroughly described, illustrated and key to their identification is provided. Species level keys are provided for Old Wold Dascillus, Petalon and Sinocaulus. Eighteen new species are described: Dascillus acutus (China: Shanxi), D. chifengi (China: Taiwan), D. compressus (Nepal), D. formosanus (China: Taiwan), D. lanceus (China: Taiwan), D. largus (China: Hubei), D. montanus (China: Sichuan), D. planus (China: Yunnan), D. russus (India: Arunachal Pradesh), D. tibetensis (China: Tibet), D. transversus (China: Yunnan), Petalon acerbas (China-Yunnan), P. allochroides (Malaysia, Sabah), P. annamensis (Vietnam), P. digitatus (China: Yunnan), P. iviei (China: Yunnan), Sinocaulus clypeatus (China: Guizhou) and S. omiensis (China: Sichuan). The following species level new synonyms are proposed (senior synonym listed first): Dascillus congruus Pascoe, 1860 (= Dascillus perroudi Pic, 1939,71. klapperichi, Pic, 1955, D. taiwanus Nakane, 1995 and D. fortunei Pic, 1913); Dascillus calvescens Bourgeois, 1892 (= D. hoizi Pic, 1911, D. rufocinctus Pic, 1913, D. brevesulcatus Pic, 1933 and A rubropubens Pic, 1934); Dascillus renardi Bourgeois, 1891(= D. rufovillosus Bourgeois, 1892); Dascillus fulvulus (Wiedemann, 1819) (= D. striatus Pic, 1911, D. corporaali Pic, 1923 and D. obscuricolor Pic, 1933); D. obscuripes Pic, 1912 (= D. rufus Pic, 1923); Pseudolichas nivipictus Fairmaire, 1904 (= Pseudolichas ruficomis Pic, 1914) and Sinocaulus rubrovelutinus Fairmaire, 1878 (= Haematoides atriceps Pic, 1910: 45). Four taxa are transferred to Dascillus Latreille (Cladotoma vittata Pic, 1914; Pseudolichas nigronotatus Pic, 1914; Pseudolichas nivipictus Fairmaire, 1904; Pseudolichas superbus Pic, 1907 and Therius jaspideus Fairmaire, 1878) comb. nov. Fifeteen species are transferred to Petalon Schonherr (Dascillus bengalensis Pic, 1911; D. birmanicus Pic, 1913; A calvescens Bourgeois, 1892; A fruhstorferi Pic, 1912; A fulvithorax Pic, 1933; A. indicus Guérin-Méneville, 1861; D. leopoldi Pic, 1933; D. major Pic, 1933; D. obscuripes Pic, 1912; D. pruinosus Fairmaire, 1896; D. renardi Bourgeois, 1891; D. rufithorax Pic, 1912; D. rufovillosus Bourgeois, 1892; D. rufus Pic, 1923 and Pseudolichas sulcifrons Fairmaire, 1878) comb. nov. The neotypes are designated for: Bruchus fulvulus Wiedemann, 1819 (Indonesia: Java) and Dascillus maculosus Fairmaire, 1889 (China: Sichuan). Lectotypes are designated for:Dascillus bengalensis Pic, 1911; D. brevesulcatus Pic, 1933; D. calvescens Bourgeois, 1892; D. cavaleriei Pic, 1930; D. corporaali Pic, 1923; D. costatus Pic, 1927; D. fortunei Pic 1913; D. holzi Pic, 1911; D. klapperichi Pic, 1955; D. nigripennis Guérin-Méneville, 1861; D. obscuricolor Pic, 1933; D. obscuripes Pic, 1912; D. pallidofemoratus Pic, 1911; D. perroudi Pic, 1939; D. renardi Bourgeois, 1891; D. rubropubens Pic, 1934; D. rufocinctus Pic, 1913; D. rufovillosus Bourgeois, 1892; D. rufus Pic, 1923; D. striatus Pic, 1911; D. sublineatus Pic, 1915; Haematoides atriceps Pic, 1910; Pseudolichas sulcifrons Fairmaire, 1878 and Sinocaulus rubrovelutinus Fairmaire, 1878.
Five species of Dicranolaius Champion: D. armstrongianus Wittmer, 1954; D. bellulm (Boisduval, 1835); D. rugulipennis (Fairmaire, 1877); D. spinicornis Wittmer, 1954 and D. villosus (Lea, 1899) are diagnosed, described and illustrated. The lectotypes of Laius rugulipennis Fairmaire, 1877 and Laius nodicornis Blackburn, 1888 have been designated.
The neotenic genus Atelius Waterhouse, 1878 from China is reviewed and two new species are described: A. brevicornis sp. nov. and A. kadoorieorum sp. nov., both from the tropical rainforests of Yunnan. The general appearance of all Chinese species and the morphology of male genitalia âre illustrated and a key to all Atelius species is provided. The distribution of Atelius and other widespread neotenic net-winged beetles in the northern part of the Oriental region indicates their survival in disjunctive regions, which were not affected by aridization and cooling of the climate. The current distribution is hypothesized as a relict of the earlier continuous range of neotenic beetles in the Oriental region.
The members of the Australian genus Australoneda Iablokoff-Khnzorian, 1984 are revised. Seven species from New Guinea and single species from Australia are recognised, described and illustrated. Two new species: A. bielawskii sp. nov. (Irian Jaya), A. ruitong sp. nov. (Papua New Guinea) are described. A key to the species is also provided.
Paradermestes jurassiens gen. et sp. nov., a new dermestid beetle is described based on a well-preserved impression fossil collected from the Middle Jurassic Jiulongshan Formation in Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China, making it the oldest fossil of the family. Based on its size, no visible ocellus and laterally reduced hind coxal plates Paradermestes is placed in subfamily Dermestinae and tribe Paradermestini, trib. nov.
The Australian members of the Coccinellid tribe Telsimini are revised which now includes 2 genera and 16 species. Nomenclatural history, diagnoses and distribution are provided for each genus and species. Keys to the genera and species are also presented. The lectotypes are designated for: Lipernes creber Blackburn, 1895, L. gibbosus Blackburn, 1895, L. subviridis Blackburn, 1892 and Serangium obscuripes Lea, 1902. Telsimia abdita, T. acaciae, T. cassicula, T. glorious, T. leucoceps, T. occidua, T. palitans, T. rossi and T. rotunda spp. nov. are described from Australia. Serangium obscuripes Lea, 1902 is transferred to Telsimia and T. elainae Chazeau is newly recorded from Australia.
A new bark-gnawing beetle genus Sinopeltis gen. nov., with two species (S. jurrasica sp. nov. (type species) and Sinopeltis amoena sp. nov.), is described based on two well-preserved impression fossils. Specimens were collected from the Middle Jurassic Jiulongshan Formation in Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China from a lacustrine paleoenvironment, making it the oldest fossil of the family. Sinopeltis is placed into Peltinae incertae sedis.
A new genus of Prionoceridae, Idgiaites gen. nov. (type species: I. jurassicus sp. nov.), is described and illustrated from a well-preserved impression fossil from the Middle Jurassic Jiulongshan Formation collected at Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China. This is the first representative of Prionoceridae described from the Mesozoic era, and the presence of this family in the Jurassic indicates that cleroids were quite diverse during the early evolution of polyphagan beetles. The occurrence of plants in the same Formation may indicate that Idgiaites like it’s modem confamilials were pollen-feeding.A new genus of Prionoceridae, Idgiaites gen. nov. (type species: I. jurassicus sp. nov.), is described and illustrated from a well-preserved impression fossil from the Middle Jurassic Jiulongshan Formation collected at Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China. This is the first representative of Prionoceridae described from the Mesozoic era, and the presence of this family in the Jurassic indicates that cleroids were quite diverse during the early evolution of polyphagan beetles. The occurrence of plants in the same Formation may indicate that Idgiaites like it’s modem confamilials were pollen-feeding.
Two enigmatic genera of Trogossitidae are described and illustrated from well-preserved impression fossils from the Middle Jurassic Jiulongshan Formation collected at Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China. Both new genera, Marginulatus gen. nov. (type species M. venustus sp. nov.) and Latitergum gen. nov. (type species L. glabrum sp. nov.) are placed in Trogossitidae incertae sedis because they have a mixture of characters that do not allow for them to be easily placed in the current classification of Trogossitidae.
A new genus of Thanerocleridae, Archaeozenodosus bellus Yu and Kolibac gen. et sp. nov., is described and illustrated from the Upper Cretaceous (Cenomanian, ca. 99 Ma) amber near the Hukawng Valley of northern Myanmar, making it the first Mesozoic record of the family. Morphological characters preserved in the Burmese amber highlight the diversity of thaneroclerids during the Late Mesozoic and provide data for future phylogenetic studies of Thanerocleridae.
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