Late Oligocene sharks and rays from the Chandler Bridge Formation, Dorchester County, South Carolina, USA

• Autorzy: Cicimurri D.J. , Knight, J.L.
• Języki publikacji: EN

Abstrakty

EN

A diverse vertebrate fauna, dominated by elasmobranch taxa, was collected from the upper Oligocene (Chattian) Chandler Bridge Formation in Summerville, Dorchester County, South Carolina. Nearly 3,500 teeth and dermal denticles are assigned to 29 species of sharks and rays, and our sample includes the oldest known occurrence of the whale shark, Rhincodon, as well as a new skate, Raja mccollumi sp. nov. The Chandler Bridge elasmobranch assemblage is comparable in species diversity to Chattian assemblages of Virginia and North Carolina, USA, and Germany. Notable absences from Germany include Rhincodon, Hemipristis, and Sphyrna zygaena, likely reflecting the influence of colder water on the North Sea Basin during the Chattian. Squaloids, pristiophoroids, and hexanchoids are known from Chattian deposits of the Albemarle Embayment (North Carolina), Salisbury Embayment (Virginia), and North Sea Basin, but these taxa are absent from the Chandler Bridge assemblage, perhaps because of shallow, warm water (20 to 25°C) conditions within the more southerly Charleston Embayment.

Słowa kluczowe

EN

Late Oligocene; Oligocene; shark; ray; Chandler Bridge Formation; Dorchester County; South Carolina; USA; Chondrichthyes; Neoselachii

• Wydawca: -
• Czasopismo: Acta Palaeontologica Polonica
• Rocznik: 2009
• Tom: 54
• Numer: 4
• Opis fizyczny: p.627-647,fig.,ref.

Twórcy

autor

Cicimurri D.J.

• Campbell Geology Museum, 140 Discovery Lane, Clemson, South Carolina 29634, USA

autor

Knight, J.L.

Bibliografia

• Adnet, S., Antoine, P.−O., Hassan Baqri, S.R., Crochet, J.−Y., Marivaux, L., Welcomme, J.−L., and Métais, G. 2007. New tropical carcharhinids (Chondrichthyes, Carcharhiniformes) from the late Eocene–early Oligocene of Balochistan, Pakistan: paleoenvironmental and paleogeographic implications. Journal of Asian Earth Sciences 30: 303–323. doi:10.1016/j.jseaes.2006.10.002
• Allen, T.B. 1999. The Shark Almanac. 274 pp. Lyons Press, New York.
• Antunes, M.T. and Balbino, A.C. 2007. Rajiformes (Neoselachii, Batomorphii) from the Alvalade basin, Portugal. Annales de Paléontologie 93: 107–119. doi:10.1016/j.annpal.2007.03.002
• Applegate, S.P. 1978. Phyletic studies, Part 1: Tiger sharks. Revista Universidad Nacional Autónoma de México, Instituto de Geología 2 (1): 55–64.
• Applegate, S.P. 1992. The case for dignathic heterodonty in fossil specimens of the tiger shark genus Galeocerdo, particularly those belonging to G. aduncus lineage. Journal of Vertebrate Paleontology 12 (Supplement to No 3): 16A.
• Applegate, S.P. and Uyeno, T. 1968. The first discovery of a fossil tooth belonging to the shark genus Heptranchias, with a new Pristiophorus spine, both from the Oligocene of Japan. Bulletin of the National Science Museum (Tokyo) 11 (2): 195–199.
• Baut, J.−P. and Génault, B. 1999. Les élasmobranches des Sables de Kerniel (Rupélien), à Gellik, nord est de la Belgique. Memoirs of the Geological Survey of Belgium 45: 1–61.
• Bigelow, H.P. and Schroeder, W.C. 1953. Fishes of the western North Atlantic, Part 2. Batoidea and Holocephali. Memoirs of the Sears Foundation of Marine Research 2: 1–562.
• Bonfil, R. and Abdallah, M. 2004. Field Identification Guide to the Sharks and Rays of the Red Sea and Gulf of Aden. 71 pp. FAO, Rome, Italy.
• Bor, T.J. 1980. Elasmobranchii from the Atuatuca Formation (Oligocene) in Belgium. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 17: 3–16.
• Bor, T.J. 1990. A new species of mobulid ray (Elasmobranchii, Mobulidae) from the Oligocene of Belgium. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 27: 93–97.
• Bosch, M. van den 1980. Elasmobranch associations in Tertiary and Quaternary deposits of the Netherlands (Vertebrata, Pisces), 2. Paleogene of the eastern and northern part of the Netherlands, Neogene in the eastern part of the Netherlands. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 17: 65–70.
• Bosch, M. van den 1981. Elasmobranchii from limonitic sandstone of Siadło Górne near Szczecin, Poland. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 18: 127–131.
• Bosch, M. van den 1984. Oligocene to Recent Cetorhinidae (Vertebrata, basking sharks); problematical finds of teeth, dermal scales, and gill rakers. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 21: 205–232.
• Bosellini, F.R. and Perrin, C. 2008. Estimating Mediterranean Oligocene–Miocene sea−surface temperatures: An approach based on coral taxonomic richness. Palaeogeography, Palaeoclimatology, Palaeoecology 258: 71–88. doi:10.1016/j.palaeo.2007.10.028
• Bracher, H. 2005. Untermiozäne Haie und Rochen. 183 pp. Privately published, Altheim.
• Brzobohatý, R. and Kalabis, V. 1970. Die Fischzähne aus Pouzdřany (Pouzdřany−Schichten, Oligozän).Acta Musei Moraviae, Scientiae Naturals 55: 41–50.
• Compagno, L.J.V. and Roberts, T.R. 1982. Freshwater stingrays (Dasyatidae) of Southeast Asia and New Guinea with description of a new species of Himantura and reports of unidentified species. Environmental Biology of Fishes 7 (4): 321–339.
• Cape Roberts Science Team 2000. Studies from the Cape Roberts Project, Ross Sea, Antarctica. Initial Report on CRP−3, Chapter 5: Paleontology. Terra Antarctica 7 (1/2): 133–170.
• Cappetta, H. 1970. Les sélaciens du Miocène de la région de Montpellier. Palaeovertebrata, Mémoire Extraordinaire 1970: 1–139.
• Cappetta, H. 1987. Chondrichthyes II. Mesozoic and Cenozoic Elasmobranchii. 193 pp. Gustav Fischer Verlag, Stuttgart, Germany.
• Cappetta, H. and Stringer, G. 2002. A new batoid genus (Neoselachii: Myliobatiformes) from the Yazoo Clay (late Eocene) of Louisiana, USA. Tertiary Research 21: 51–56.
• Case, G.R. 1980. A selachian fauna from the Trent Formation, lower Miocene (Aquitanian) of eastern North Carolina. Palaeontographica A 171: 75–103.
• Case, G.R. 1981. Late Eocene selachians from south central Georgia. Palaeontographica A 176: 52–79.
• Chandler, R.E., Chiswell, K.E., and Faulkner, G.D. 2006. Quantifying a possible Miocene phyletic change in Hemipristis (Chondrichthyes) teeth, 14 p. Palaeontologica Electronica http://palaeo−electronica.org/2006_1/teeth/teeth.pdf.
• Clandenin, C.W., Willoughby, R.H., and Niewendorp, C.A. 1999. The Coastal Plain region of South Carolina: A review of tectonic influences and feedback processes. South Carolina Geology 41: 45–63.
• Compagno, L.J.V. 1984. FAO Species Catalogue. Volume 4. Sharks of the World. 249 pp. (part 1) and 655 pp. (part 2). FAO, Rome, Italy.
• Compagno, L.J.V. and Roberts, T.R. 1982. Freshwater stingrays (Dasyatidae) of Southeast Asia and New Guinea with description of a new species of Himantura and reports of unidentified species. Environmental Biology of Fishes 7 (4): 321–339.
• Compagno, L.J.V., Dando, M., and Fowler, S. 2005. Sharks of the World. 368 pp. Princeton University Press, Princeton, New Jersey.
• De Man, E. 2003. Foraminiferal biofacies analysis of the Rupelian–Chattian transition in the Weelde borehole (northern Belgium). Geologica Belgica 6 (3–4): 201–211.
• De Man, E., and Van Simaeys, S. 2004. Late Oligocene warming event in the southern North Sea Basin: benthic foraminifera as paleotemperature proxies. Netherlands Journal of Geosciences 83: 227–239.
• Dockery, D.T. III and Lozouet, P. 2003. Molluscan faunas across the Eocene/Oligocene boundary in the North American Gulf Coastal Plain, with comparisons to those of the Eocene and Oligocene of France. In: D.R. Prothero, L.C. Ivany, and E.A. Nesbitt (eds.), From Greenhouse to Icehouse: The Marine Eocene–Oligocene Transition, 303–340. Columbia University Press, New York.
• Dockery, D.T. III and Manning, E.M. 1986. Teeth of the giant shark Carcharodon auriculatus from the Eocene and Oligocene of Mississippi. Mississippi Geology 7: 7–19.
• Edwards, L.E., Bybell, L.M., Gohn, G.S., and Frederiksen, N.O. 1997. Paleontology and physical stratigraphy of the USGS−Pregnall No. 1 core (DOR−208), Dorchester County, South Carolina. USGS Open−file Report 97−145: 1–35.
• Edwards, L.E., Gohn, G.S., Bybell, L.M., Chirico, P.G., Christopher, R.A., Frederiksen, N.O., Prowell, D.C., Self−Trail, J.M., and Weems, R.E. 2000. Supplement to the preliminary stratigraphic database for subsurface sediments of Dorchester County, South Carolina. USGS Open−file Report 00−049−B: 1–44.
• Elewa, A.M.T. 2002. Paleobiogeography of Maastrichtian to early Eocene Ostracoda of North and West Africa and the Middle East. Micropaleontology 48 (4): 391–398.
• Erickson, B.R. 1990. Paleoecology of crocodile and whale−bearing strata of Oligocene age in North America. Historical Biology 4: 1–14. doi:10.1080/08912969009386530
• Fierstine, H.L. 2001. A new Aglyptorhynchus (Perciformes: Scombroidei: †Blochiidae) from the late Oligocene of Oregon. Journal of Vertebrate Paleontology 21: 24–33. doi:10.1671/0272−4634(2001)021[0024:ANAPSB] 2.0.CO;2
• Fierstine, H.L. 2005. A new Aglyptorhynchus (Perciformes: Scombroidei) from the Lincoln Creek Formation (late Oligocene, Washington, U.S.A.). Journal of Vertebrate Paleontology 25 (2): 288–299. doi:10.1671/0272−4634(2005)025[0288:ANAPSF]2.0.CO;2
• Flower, B.P. and Chisholm, K.E. 2006. Magnetostratigraphic calibration of the late Oligocene climate transition. In: R. Tiedemann, A.C. Mix, R. Richter, and W.F. Ruddiman (eds.), Proceedings of the Ocean Drilling Program, Scientific Results 202: 1–15.
• Freile, D., DeVore, M.L., and Parmley, D. 2001. The first record of Carcharocles auriculatusfrom the Oligocene of Georgia in the context of previous Gulf Coast records. Georgia Journal of Science 59 (3): 128–136.
• Génault, B. 1993. Contribution à l’étude des élasmobranches Oligocènes du bassin de Paris 2. Découverte de deux horizons à élasmobranches dans le Stampien (Sables de Fontainebleau) de la feuille géologique de Chartres. Cossmanniana 2: 13–36.
• González−Isáis, M. and Domínguez, H.M.M. 2004. Comparative anatomy of the Superfamily Myliobatoidea (Chondrichthyes) with some comments on phylogeny. Journal of Morphology 262: 517–535. doi:10.1002/jmor.10260
• Gottfried, M.D. 1993. An associated tiger shark dentition from the Miocene of Maryland. The Mosasaur 5: 59–61.
• Gottfried, M.D. and Fordyce, R.E. 2001. An associated specimen of Carcharodon angustidens (Chondrichthyes, Lamnidae) from the late Oligocene of New Zealand, with comments on Carcharodon interrelationships. Journal of Vertebrate Paleontology 28: 730–739. doi:10.1671/0272−4634(2001)021[0730:AASOCA]2.0.CO;2
• Gottfried, M.D., Rabarison, J.A., and Randriamiarimanana, L.L. 2001. Late Cretaceous elasmobranchs from the Mahajanga Basin of Madagascar. Cretaceous Research 22: 491–496.
• Hankins, K.G. and Prothero, D.R. 2001. Magnetic stratigraphy and tectonic rotation of the Eocene–Oligocene Keasey and Pittsburg Bluff formations, northwestern Oregon. Abstracts with programs, joint meeting of the Cordilleran Section of the Geological Society of America and Pacific Section of the American Association of Petroleum Geologists, April 9–11, 2001. Universal City, California.
• Harris, W.B. and Zullo, V.A. 1991. Eocene and Oligocene stratigraphy of the outer Coastal Plain.In: J.W. Horton and V.A. Zullo (eds.), The Geology of the Carolinas, 251–262. University of Tennessee Press, Knoxville.
• Harris, W.B., Mendrick, S., and Fullagar, P.D. 2000. Correlation of onshore−offshore Oligocene through lower Miocene strata using 87Sr/86Sr isotopic ratios, north flank of Cape Fear Arch, North Carolina, USA. Sedimentary Geology 134: 49–63. doi:10.1016/S0037−0738(00)00013−0
• Haye, T., Reinecke, T., Gürs, K., and Piehl, A. 2008. Die Elasmobranchier des Neochattiums (Oberoligozän) von Johannistal, Ostholstein, und Ergänzungen zu deren vorkommen in der Ratzeburg−Formation (Neochattium) des Südöstlichen Nordseebeckens. Palaeontos 14: 55–95.
• Hazin, F.H.V., Vaske Junior, T., Oliveira, P.G., Macena, B.C.L., and Carvalho, F. 2008. Occurences of the whale shark (Rhincodon typus Smith, 1829) in the Saint Peter and Saint Paul archipelago, Brazil. Brazilian Journal of Biology 68 (2): 385–389. doi:10.1590/S1519−69842008000200021
• Herman, J. 1975. Quelques restes de sélaciens récoltés dans les Sables du Kattendijk à Kallo. 1 Selachii – Euselachii. Bulletin de la Société belge de Géologie 83: 15–31.
• Herman, J., Hovestadt−Euler, M., and Hovestadt, D.C. 1988. Part A: Selachii. N°2a: Order: Carcharhiniformes—Family: Triakidae. In: M. Stehmann (ed.), Contributions to the study of the comparative morphology of teeth and other relevant ichthyodorulites in living supraspecific tax of Chondrichthyan fishes. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 58: 99–126.
• Herman, J., Hovestadt−Euler, M., and Hovestadt, D.C. 1990. Part A: Selachii, N°2b: Order: Carcharhiniformes—Family: Scyliorhinidae. In: M. Stehmann (ed.), Contributions to the study of the comparative morphology of teeth and other relevant ichthyodorulites in living supraspecific tax of Chondrichthyan fishes. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 60: 181–230.
• Herman, J., Hovestadt−Euler, M., and Hovestadt, D.C. 1992. Part A: Selachii, N°4: Order: Orectolobiformes. Families: Brachaeluridae, Ginglymostomatidae, Hemiscylliidae, Orectolobidae, Parascylliidae, Rhiniodontidae, Stegostomatidae. Order: Pristiophoriformes—Family: Pristiophoridae. Order: Squatiniformes—Family: Squatinidae. In: M. Stehmann (ed.), Contributions to the study of the comparative morphology of teeth and other relevant ichthyodorulites in living supraspecific tax of Chondrichthyan fishes. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 62: 193–254.
• Herman, J., Hovestadt−Euler, M., and Hovestadt, D.C. 1993. Part A: Selachii, N°1b: Order: Hexanchiformes—Family: Chlamydoselachidae; N°5: Order: Heterodontiformes—Family: Heterodontidae; N°6: Order: Lamniformes—Families: Cetorhinidae, Megachasmidae; Addendum 1 to N°3: Order Squaliformes; Addendum 1 to N°4: Order: Orectolobiformes; General Glossary; Summary Part A. In: M. Stehmann (ed.), Contributions to the study of the comparative morphology of teeth and other relevant ichthyodorulites in living supraspecific tax of Chondrichthyan fishes. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 63: 185–256.
• Hocht, F. von der. 1978a. Bestandsaufnahme der Chondrichthyes−Fauna des unteren Meeressandes (Oligozän, Rupelium) im Mainzer Becken. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 15: 77–83.
• Hocht, F. von der. 1978b. Verbreitung von Chondrichthyes−Arten (Vertebrata, Pisces) im Rupelium des Mainzer Beckens und im Chattium von Norddeutschland. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie 15: 163–165.
• Hovestadt, D.C. and Hovestadt−Euler, M. 1995. Additions to the fauna of the Boom Clay Formation of Belgium (Rupelian, Oligocene). Taxonomic adjustments on the Scyliorhinidae and Rajoidei, discovery of a dasyatid species (Pisces, Chondrichthyes) and of a curculionid species (Insecta, Coleoptera). Belgian Geological Survey Professional Paper 278: 261–282.
• Hovestadt, D.C. and Hovestadt−Euler, M. 1999. Weissobatis micklichi n. gen., n. sp., an eagle ray (Myliobatiformes, Myliobatidae) from the Oligocene of Frauenweiler (Baden−Wurttemberg, Germany). Paläontologische Zeitschrift 73 (3/4): 337–349.
• Kajiura, S.M. and Tricas, T.C. 1996. Seasonal dynamics of dental sexual dimorphism in the Atlantic stingray Dasyatis sabina. Journal of Experimental Biology 199: 2297–2306.
• Kajiura, S.M., Sebastian, A.P., and Tricas, T.C. 2000. Dermal bite wounds as indicators of reproductive seasonality and behaviour in the Atlantic Stingray, Dasyatis sabina. Environmental Biology of Fishes 58: 23–31. doi:10.1023/A:1007667108362
• Katuna, M.P., Geisler, J.H., and Colquhoun, D.J. 1997. Stratigraphic correlation of Oligocene marginal marine and fluvial deposits across the middle and lower coastal plain, South Carolina. Sedimentary Geology 108: 181–194. doi:10.1016/S0037−0738(96)00053−X
• Kemp, N.R. 1982. Chondrichthyes in the Tertiary of Australia. In: P.V. Rich and E.M. Thompson (eds.), The Fossil Vertebrate Record of Australasia, 498–531. Monash University, Clayton, Victoria, Australia.
• Kent, B.W. 1994. Fossil Sharks of the Chesapeake Bay Region. 146 pp. Eegan Rees and Boyer Inc., Columbia, Maryland.
• Keyes, I.W. 1979. Ikamauius, a new genus of fossil sawshark (Order Selachii: Family Pristiophoridae) from the Cenozoic of New Zealand. New Zealand Journal of Geology and Geophysics 22: 125–129.
• Keyes, I.W. 1982. The Cenozoic sawshark Pristiophorus lanceolatus (Davis) (Order Selachii) of New Zealand and Australia, with a review of the phylogeny and distribution of world fossil and extant Pristiophoridae. New Zealand Journal of Geology and Geophysics 25: 459–474.
• Kier, P.M. 1997. Oligocene echinoids from North Carolina. Smithsonian Contributions to Paleobiology 83: 1–37.
• Kruckow, T. and Thies, D. 1990. Die Neoselachier der Palaokaribik (Pisces: Elasmobranchii).Courier Forschungsinstitut Senckenberg 119: 1–102.
• Leriche, M. 1908. Note préliminaire sur les poissons nouveaux de l’Oligocène belge. Bulletin de la Société belge de Géologie, de Paléontologie et d’Hydrologie 22: 1–379.
• Leriche, M. 1910. Les poissons Oligocènes de la Belgique. Mémoires du Musée Royal d’Histoire Naturelle de Belgique 5: 233–363.
• Leriche, M. 1927. Les poissons de la Molasse Suisse. Mémoires de la Société Paléontologique Suisse 46: 1–119.
• Leriche, M. 1942. Contribution à l’étude des faunes ichthyologiques marines des terrains tertiares de la plaine cotière Atlantique et du centre des États−Unis. Mémoires de la Société Géologique de France (nouvelle série), Mémoire 20 (45): 1–110.
• Manning, E.M. 2006. The Eocene/Oligocene transition in marine vertebrates of the Gulf Coastal Plain. In: D.R. Prothero, L.C. Ivany, and E.A. Nesbitt (eds.), From Greenhouse to Icehouse: The Marine Eocene–Oligocene Transition, 366–385. Columbia University Press, New York.
• Marsili, S., Carnevale, G., Danese, E., Bianucci, G., and Landini, W. 2007. Early Miocene vertebrates from Montagna della Maiella, Italy. Annales de Paléontologie 93: 27–66. doi:10.1016/j.annpal.2007.01.001
• Miller, K.G., McLaughlin, P.P., Browning, J.V., Benson, R.N., Sugarman, P.J., Hernandez, J., Ramsey, K.W., Baxter, S.J., Feigenson, M.D., Aubry, M.−P., Monteverde, D.H., Cramer, B.S., Katz, M.E., McKenna, T.E., Strohmeier, S.A., Pekar, S.F., Uptegrove, J., Cobbs, G., Cobbs, G.III, and Curtin, S.E. 1999. Bethany Beach site report, ODP Leg 174AX. Proceedings of the Ocean Drilling Program, Initial Reports 174AX, 1–85. (published online at www−odp.tamu.edu/publications/174AXSIR/chap_03/chap_03.htm)
• Miller, M.M. 2000. Paleoecology of the Oligocene Mint Spring Formation Based on Otoliths and Related Vertebrates. 54 pp. Unpublished MSc. thesis. The University of Louisiana, Monroe.
• Monkolprasit, S. and Roberts, T.R. 1990. Himantura chaophraya, a new giant freshwater stingray from Thailand. Japanese Journal of Ichthyology 37: 203–208.
• Müller, A. 1983. Fauna und Palökologie des marinen Mitteloligozäns der Leipziger Tieflandsbucht (Böhlener Schichten). Altenburger Naturwissenschaftliche Forschungen 2: 1–152.
• Müller, A. 1999. Ichthyofaunen aus dem atlantischen Tertiär der USA. Leipziger Geowissenschaften 9/10: 1–360.
• Notabartolo di Sciara, G. 1987. A revisionary study of the genus Mobula Rafinesque, 1810 (Chondrichthyes: Mobulidae) with the description of a new species. Zoological Journal of the Linnean Society 91: 1–91. doi:10.1111/j.1096−3642.1987.tb01723.x
• Odin, G.S. and Fullagar, P.D. 1988. Geological significance of the glaucony facies. In: G.S. Odin (ed.), Green Marine Clays, 295–332. Elsevier, Amsterdam.
• Parmley, D. and Cicimurri, D.J. 2003. Late Eocene sharks of the Hardie Mine local fauna of Wilkinson County, Georgia. Georgia Journal of Science 61: 153–178.
• Pekar, S.F., DeConto, R.M., and Harwood, D.M. 2006. Resolving a late Oligocene conundrum: Deep−sea warming and Antarctic glaciation. Palaeogeography, Palaeoclimatology, Palaeoecology 231: 29–40. doi: 10.1016/j.palaeo.2005.07.024
• Pfeil, F.H. 1981. Eine nektonische Fischfauna aus dem unteroligozänen Schönecker Fischschiefer des Galon−Grabens in Oberbayern. Geologica Bavarica 82: 357–388.
• Pfeil, F.H. 1984. Neoselachian teeth collected from phosphorite−bearing greensand on Chatham Rise east of New Zealand. Geologisches Jahrbuch 65: 107–115.
• Pledge, N.S. 1967. Fossil elasmobranch teeth of South Australia and their stratigraphic distribution. Transactions of the Royal Society of South Australia 91: 135–160.
• Purdy, R.W. 1996. Paleoecology of fossil great white sharks. In: A.P. Klimley and D. Ainley (eds.), The Biology of the White Shark, Carcharodon carcharias. 517 pp. Academic Press, San Diego, California, USA.
• Purdy, R.W. 1998. The early Miocene fauna from the Pollack Farm site, Delaware. In: R.N. Benson (ed.), Geology and paleontology of the lower Miocene Pollack Farm Fossil site, Delaware. Delaware Geological Survey Special Publication 21: 133–139.
• Purdy, R.W., Schneider, V.P., Applegate, S.P., McLellan, J.H., Meyer, R.L., and Slaughter, B. H. 2001. The Neogene sharks, rays and bony fishes from Lee Creek Mine, Aurora, North Carolina. Smithsonian Contribution to Paleontology 90: 71–202.
• Reinecke, T. and Hoedemakers, K. 2006. Physogaleus hemmooriensis (Carcharhinidae, Elasmobranchii) a new shark species from the early to middle Miocene of the North Sea Basin. Palaeovertebrata 34: 1–25.
• Reinecke, T., Hocht, F. von der, and Gürs, K. 2008. Die Elasmobranchier des Vierlandiums, Unteres Miozän, im Nordwestdeutschen Becken aus Bohrungen und glaziofluvialiten Geröllen (“Holsteiner Gestein”) der Vierlande−Feinsande (Holstein) und der Kakert−Schichten (Niederrhein). Palaeontos 14: 1–54.
• Reinecke, T., Moths, H., Grant, A., and Breitkreutz, H. 2005. Die Elasmobranchier des Norddeutschen Chattiums, insbesondere des Sternberger Gesteins (Eochattium, Oberes Oligozän). Palaeontos 8: 1–135.
• Reinecke, T., Stapf, H., and Raisch, M. 2001. Die Selachier und Chimären des Unteren Meeressandes und Schleichsandes im Mainzer Becken (Rupelium, unteres Oligozän). Palaeontos 1: 1–73.
• Rossbach, T.J., and Carter, J.G. 1991. Molluscan biostratigraphy of the lower River Bend Formation at the Martin Marietta quarry, New Bern, North Carolina. Journal of Paleontology 65: 80–118.
• Sanders, A.E. 1980. Excavation of Oligocene marine fossil beds near Charleston, South Carolina.National Geographic Society Research Report 12: 601–621.
• Sanders, A.E. 2002. Additions to the Pleistocene mammal faunas of South Carolina, North Carolina, and Georgia. Transactions of the American Philosophical Society 92: 1–152. doi:10.2307/4144916
• Sanders, A.E., Weems, R.E., and Lemon, E.M. Jr. 1982. Chandler Bridge Formation—a new Oligocene stratigraphic unit in the lower coastal plain of South Carolina. United States Geological Survey Bulletin 1529−H: 105–124.
• Segall, M.P., Siron, D.L., and Colquhoun, D.J. 2000. Depositional and diagenetic signatures of late Eocene–Oligocene sediments, South Carolina. Sedimentary Geology 134: 27–47. doi:10.1016/S0037−0738(00)00012−9
• Snyder, S.W., Müller, C., and Miller, K.G. 1983. Biostratigraphy and paleoeoceanography across the Eocene/Oligocene boundary at Deep Sea Drilling Project Site 549. Initial Reports of the Deep Sea Drilling Project 80: 567–572.
• Southall, E.J. and Sims, D.W. 2005. A smooth hammerhead shark (Sphyrna zygaena) from south−west England. JMBA2—Biodiversity Records, 1–2. Marine Biological Association of the United Kingdom. (published online at www.mba.ac.uk/jmba/pdf/5098.pdf)
• Steurbaut, E. and Herman, J. 1978. Biostratigraphie et poissons fossiles de la Formation de L’Argile de Boom (Oligocène Moyen du Bassin Belge). Geobios 11: 297–325. doi:10.1016/S0016−6995(78)80033−3
• Stolarski, J. and Taviani, M. 2001. Oligocene scleractinian corals from CRP−3 drillhole, McMurdo Sound (Victoria Land Basin, Antarctica). Terra Antarctica 8 (3): 1–4.
• Stringer, G.L., Beard, S.Q., and Kontrovitz, M. 2001. Biostratigraphy and paleoecology of diagnostic invertebrates and vertebrates from the type locality of the Oligocene Rosefield Marl Beds, Louisiana. Gulf Coast Association of Geological Societies Transactions 51: 321–328.
• Thomas, H., Roger, J., Sen, S., Bourdillon−de−Grissac, C., and al−Sulaimani, Z. 1989. Découverte de vértébres fossiles dans l’Oligocène inférieur du Dhofar (Sultanat d’Oman). Geobios 22: 101–120. doi:10.1016/S0016−6995 (89)80091−9
• Uyeno, T., Yabumoto, Y., and Kuga, N. 1984. Fossil fishes of the Ashiya Group—(I) Late Oligocene elasmobranchs from the islands of Ainoshima and Kaijima, Kitakyushu [in Japanese, with English abstract]. Bulletin of the Kitakyushu Museum of Natural History 5: 135–142.
• Van Semaeys, S. and Vandenberghe, E. 2006. Rupelian. Geologica Belgica 9 (1–2): 95–101.
• Van Semaeys, S., De Man, E., Vandenberghe, N., Brinkhuis, H., and Steurbaut, E. 2004. Stratigraphic and palaeoenvironmental analysis of the Rupelian–Chattian transition in the type region: evidence from dinoflagellate cysts, foraminifer and calcareous nannofossils. Palaeogeography, Palaeoclimatology, Palaeoecology 208: 31–58. doi:10.1016/j.palaeo.2004.02.029
• Ward, L.W. 1992. Tertiary molluscan assemblages from the Salisbury Embayment of Virginia.Virginia Journal of Science 43 (1B): 85–100.
• Ward, D.J. and Bonavia, C.G. 2001. Additions to, and a review of, the Miocene shark and ray fauna of Malta. The Central Mediterranean Naturalist 3 (3): 131–146.
• Weems, R.E. and Sanders, A.E. 1986. The Chandler Bridge Formation (upper Oligocene) in the Charleston region, South Carolina. Geological Society of America Centennial Field Guide (Southeastern Section) 6: 323–326. doi:10.1130/0−8137−5406−2.323
• Welton, B.J. 1972. Fossil sharks in Oregon. The Ore Bin 34 (10): 161–170.
• Welton, B.J. 1973. Oligocene selachians from the Keasey Formation at Mist, Oregon. Abstracts with programs, Geological Society of America, Cordilleran Section Annual Meeting, 121. Protland, Oregon.
• Welton, B.J. 1979. Late Cretaceous and Cenozoic Squalomorphii of the Northwest Pacific Ocean. 553 pp. Unpublished Ph.D. thesis. University of California, Berkeley.
• Welton, B.J. and Farish, R.F. 1993. The Collector’s Guide to Fossil Sharks and Rays from the Cretaceous of Texas. 204 pp. Before Time, Louisville, Texas.
• Wienrich, G., and Reinecke, T. 2009. Elasmobranchii.In: G. Wienrich (ed.), Die Fauna des marinen Miozäns von Kevelaer (Niederrhein), 972–993. Backhuys Publishers, Leiden, Germany.
• White, E.I. 1956. The Eocene fishes of Alabama. Bulletins of American Paleontology 36 (156): 123–152.
• Yabumoto, Y. 1987. Oligocene lamnid shark of the genus Carcharodon from Kitakyushu, Japan. Bulletin of the Kitakyushu Museum of Natural History 6: 239–264.
• Yabumoto, Y. and Uyeno, T. 1994. Late Mesozoic and Cenozoic fish faunas of Japan. Island Arc 3 (4): 255–269. doi:10.1111/j.1440−1738.1994.tb00115.x