Synopsis of the terrestrial vertebrate faunas from the Middle Kura Basin (Eastern Georgia and Western Azerbaijan, South Caucasus)

• Autorzy: Bukhsianidze M. , Koiava K.
• Języki publikacji: EN

Abstrakty

EN

This paper summarizes knowledge on the Neogene–Quaternary terrestrial fossil record from the Middle Kura Basin accumulated over a century and aims to its integration into the current research. This fossil evidence is essential in understanding the evolution of the Eurasian biome, since this territory is located at the border of Eastern Mediterranean and Central Asian regions. The general biostratigraphic framework suggests existence of two major intervals of the terrestrial fossil record in the area, spanning ca. 10–7 Ma and ca. 3–1 Ma, and points to an important hiatus between the late Miocene and late Pliocene. General aspects of the paleogeographic history and fossil record suggest that the biogeographic role of the Middle Kura Basin has been changing over geological time from a refugium (Khersonian) to a full-fledged part of the Greco-Iranian province (Meotian–Pontian). The dynamic environmental changes during the Quaternary do not depict this territory as a refugium in its general sense. The greatest value of this fossil record is the potential to understand a detailed history of terrestrial life during demise of late Miocene Hominoidea in Eurasia and early Homo dispersal out of Africa. Late Miocene record of the Middle Kura Basin captures the latest stage of the Eastern Paratethys regression, and among other fossils counts the latest and the easternmost occurence of dryopithecine, Udabnopithecus garedziensis, while the almost uninterrupted fossil record of the late Pliocene–Early Pleistocene covers the time interval of the early human occupation of Caucasus and Eurasia.

• Wydawca: -
• Czasopismo: Acta Palaeontologica Polonica
• Rocznik: 2018
• Tom: 63
• Numer: 3
• Opis fizyczny: p.441-461,fig.,ref.

Twórcy

autor

Bukhsianidze M.

• Georgian National Museum, 3, Purtseladze street, Tbilisi 0105, Georgia

autor

Koiava K.

• Alexandre Janelidze Institute of Geology, A. Politkovskaia str. 31, Tbilisi 0186,vane Javakhishvili Tbilisi State University, Georgia

Bibliografia

• Adamia, S., Alania, V., Chabukiani, A., Chichua, G., Enukidze, O., and Sadradze, N. 2010. Evolution of the Late Cenozoic basins of Georgia (SW Caucasus): a review. Geological Society, London, Special Publications 340: 239–259.
• Agustí, J., Sanz de Siria, A., and Garcés, M. 2003. Explaining the end of hominoid experiment in Europe. Journal of Human Evolution 45: 145–153.
• Agustí, J., Vekua, A., Oms, O., Lordkipanidze, D., Bukhsianidze, M., Kiladze, G., and Rook, L. 2009. The Pliocene–Pleistocene succession of Kvabebi (Georgia) and the background to the early human occupation of South Caucasus. Quaternary Science Reviews 28: 3275–3280.
• Akhalkatsi, M. and Tarkhnishvili, D. 2012. Habitats of Georgia. 118 pp. WWF, Tbilisi.
• Alania, V., Chabukiani, A., Chagelishvili, R., Enukidze, O., Gogrichiani, K., Razmadze, A., and Tsereteli, N. 2016. Growth structures, piggy-back basins and growth strata of Georgian part of Kura foreland fold and thrust belt: implication for Late Alpine kinematic evolution. Geological Society, London, Special Publications 428: 171–185.
• Ali-Zade, A.A. 1961. Akčagil Turkmenistana. 300 pp. Gosgeoltehizdat, Moskva.
• Ali-Zade, A.A. 1973. Apšeron Azerbajdžana. 228 pp. Nedra, Moskva.
• Ali-Zade, A.A., Alizade, K.A., Aleskerov, D.A., Buleishvili, D.A., Vekua, A.K., Konstantinova, N.A., Lebedeva, N.A., Negadaev-Nikonov, K.N., Bikiforova, K.V., Pevzner, M.A., Khubka, A.N, Chepalyga, A.L., and Chernyakhovsky, A.G. 1972. Guidbook Excursions in Moldavia, Georgia, Azerbaijan May–June, 1972. 104 pp. INQUA Subcommission on the Pliocene/Quaternary Boundary, IUGS Subcommission on Neogene Stratigraphy, Geological Institute of the Academy of Sciences of the USSR, Department of Paleontology and Stratigrapby of the Academy of Sciences of Moldavian SSR, Institute of Paleobiology of the Academy of Sciences of Georgian SSR, Institute of Geology of the Academy of Sciences of Azerbaijan SSR, Azerbaijan Research Design Institute of Oil Prospecting.
• Alizade, K.A. 1954. Akčagil’skij Ârus Azerbajdžana. 336 pp. AS AzSSR, Baku.
• Andrianov, K.C. and Larin, N.I. 1935. Conditions of fossilization of bones of the upper Sarmatian vertebrates in the step El’dar [in Russian]. Bûlleten’ Moskovskogo Obŝestva Ispytatelej Prirody, Otdel Geologičeskij, Novaâ Seriâ 43 (4): 559–569.
• Bakradze, M.A. and Chkhikvadze, V.M. [Čhikvadze, V.M.] 1984. Fossil terrestrial turtles of the genus Testudo in the USSR [in Russian]. Vestnik Gosudarstvennogo Muzeâ Gruzii 33A: 163–174.
• Begun, D.R. 2001. African and Eurasian Miocene hominoids and the origins of the Hominidae. In: L. de Bonis, G.D. Koufos, and P. Andrews (eds.), Hominoid Evolution and Climate Change in Europe, Vol. 2: Phylogeny of the Neogene Hominoid Primates of Eurasia, 231–253. Cambridge University Press, Cambridge.
• Begun, D.R. 2010. Miocene Hominids and the Origins of the African Apes and Humans. The Annual Review of Anthropology 39: 67–84.
• Bendukidze, O.G. and Burchak-Abramovich, N.O. [Burčak-Abramovič, N.I.] 1990. Fossil beavers from the Miocene deposits of eastern Georgia [in Russian]. In: I.G. Taktakišvili (ed.), Flora i Fauna Mezo-Kainozoâ Gruzii, 145–151. Mecniereba, Tbilisi.
• Bendukidze, O.G. and Vekua, A.K. 2012. The fossil marten from Kvabebi (eastern Georgia, Kakheti) [in Russian with abstract in English]. Sak’art’velos Erovnuli Muzeumis Mac’ne Sabunebismetqvelo da Preistoriuli Sek’c’ia 4: 64–70.
• Bendukidze, O.G., Vanishvili, N. [Vanišvili, N.], and Vekua, A.K. 2013. Simocyon—a new element in the Udabno mammal fauna [in Russian with abstract in English]. Sak’art’velos Erovnuli Muzeumis Mac’ne Sabunebismetqvelo da Preistoriuli Sek’c’ia 5: 97–104.
• Bernor, R.L., Solounias, N., Swisher III, C.C., and Van Couvering, J.A. 1996. The correlation of three classical “Pikermian” mammal faunas—Maragheh, Samos, Pikermi—with the European MN Unit System. In: R.L. Bernor, V. Fahlbusch, and H.-W. Mittman (eds.), The Evolution of Western Eurasian Neogene Mammal Faunas, 137–154. Columbia University Press, New York.
• Birks, H.J.B. and Tinner, W. 2016. Past forests of Europe. In: J. San-Miguel-Ayanz, D. de Rigo, G. Caudullo, T. Houston Durrant, and A. Mauri (eds.), European Atlas of Forest Tree Species, 36–39. Publications Office of the European Union, Luxembourg.
• Bukhsianidze, M., Chagelishvili, R., and Lordkipanidze, D. 2018. Late Miocene vertebrate site of Chachuna (Iori valley, Georgia, Southern Caucasus). Bulletin of the Georgian National Academy of Sciences 12 (2): 70–75.
• Buleishvili, D.A. [Buleišvili, D.A.] 1960. Geologiâ i Neftegazenosnost’ Mežgorni Vpadiny Vostočnoj Gruzii. 238 pp. Gostoptehizdat, Leningrad.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] 1957. Fossil goose from the Udabno hipparion fauna [in Russian]. Doklady Akademii Nauk Azerbajdžanskoj SSR 13: 655–659.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Gabashvili, E.G. [Gabašvili, E.G.] 1945. A superior anthropoid ape from the Late Tertiary deposits of east Georgia [in Georgian and Russian with English abstract]. Soobŝeniâ Akademii Nauk Gruzinskoj SSR 6 (6): 451–464.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Gabashvili, E.G. [Gabašvili, E.G.] 1946. A superior antropoid from the Upper Tertiar deposits of the eastern Georgia [in Georgian and Russian with abstract in English]. Vestnik Gosudarstvennogo Muzeâ Gruzii 13A: 235–252.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Gabashvili, E.G. [Gabašvili, E.G.] 1980. Fossil beavers in the Caucasus [in Russian]. Izvestiâ Geologičeskogo Obŝestva Gruzii 6: 20–34.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Gabashvili, E.G. [Gabašvili, E.G.] 1984. Fossil gull in the hipparion fauna Garedjis Udabno [in Russian]. Vestnik Gosudarstvennogo Muzeâ Gruzii 33A: 3–13.
• Burchak-Abramovich, N.I. and Gadzyev, D.V. 1978. Anser eldaricus sp. nov. from upper Sarmatian hipparion fauna of Eldar. Acta Zoologica Cracoviensia 23: 67–68.
• Burchak-Abramovich, N.I. and Meladze, G.K. 1972. Rustaviornis georgicus gen. et sp. n., a new fossil bird from the Hipparion fauna of Georgia. Acta Zoologica Cracoviensia 17: 373–388.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Vekua, A.K. 1980. Paleobiologičeskaâ istoriâ pozdnekajnozoiskih bykov Kavkaza. 92 pp. Mecniereba, Tbilisi.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.] and Vekua, A.K. 1981. Fossil bustard-runner of East Georgian Akchagil [in Russian with abstract in English]. Soobŝeniâ Akademii Nauk Gruzinskoj SSR, Seriâ Biologičeskaâ 7 (1): 53–59.
• Burchak-Abramovich, N.I. [Burčak-Abramovič, N.I.], Gadzhiev, D.V. [Gadžiev, D.V.], and Vekua, A.K. 1980. Concerning ancestral form of bisons from the Akchagyl of South Caucasus [in Russian]. Soobŝeniâ Akademii Nauk Gruzinskoj SSR 97 (2): 485–488.
• Casanovas-Vilar, I., Alba, D.M., Garcés, M., Roblesa, J.M., and MoyàSolà, S. 2011. Updated chronology for the Miocene hominoid radiation in Western Eurasia. Proceedings of the National Academy of Sciences 108 (14): 5554–5559.
• Casanovas-Vilar I., Moyá-Solá S., Agustí J., and Köhler M. 2005. The geography of a faunal turnover: tracking the Vallesian crisis. In: A.M.T. Elewa (eds.), Migration of Organisms, 247–300. Springer, Berlin.
• Chelidze, G. 1955. [Čelidze, G.] The geological structure of the Kura Gorge between city Rustavi and the Krasnyj bridge [in Georgian, with summary in Russian]. Trudy Geologičeskogo Instituta Akademii Nauk Gruzinskoj SSR, Seriâ Geologičeskaâ 7 (13): 186–215.
• Chkhikvadze, V.M. and Amiranashvili, N.G. 1996. History of the fossil turtles studies in the Caucasus. Bulletin of the Georgian Academy of Sciences 154 (3): 470–474.
• Chkhikvadze, V.M. [Čhikvadze, V.M.] and Petrov, V.A. 2000. New late Neogene turtle localities in the Iori Valley (eastern Georgia) [in Russian]. In: N.B. Ananjeva, I. Darevskij, N.N. Dunaev, N.N. Iordanskij, S.L. Kuzmin, and N.L. Orlov (eds.), Voprosy Gerpetologii, Materialy Pervogo S”ezda Gerpetologičeskogo Obŝestva Imeni A.M. Nikol’skogo, 4–7 Dekabrâ 2000, 332–334. Puŝino, Moskva.
• Chkhikvadze, V.M. [Čhikvadze, V.M.], Mchedlidze, G.A. [Mčedlidze, G.A.], Burchak-Abramovich, N.O. [Burčak-Abramovič, N.O.], Bendukidze, O.G., Burchak, D.N. [Burčak, D.N.], Gabelaia, T.D., Amiranashvili, N.G. [Amiranašvili, N.G.], Meladze, G.K., and Kharabadze, E.S. [Harabadze, E.S.] 2000. Review of the localities of tertiary vertebrates of Georgia [in Georgian, with abstract in Russian]. Trudy Geologičeskogo Instituta Imeni A.I. Džanelidze Akademii Nauk Gruzin skoj SSR Novaâ Seriâ 115: 153–162.
• Chubinishvili, K.G. [Čubinišvili, K.G.] 1982. Neogenovye Kontinental’nye Molassy Vostočnoj Gruzii. 314 pp. Mecniereba, Tbilisi.
• Chumakov, I.S. [Čumakov, I.S.], Byzova, S.L., and Ganzei, S.S. 1992. Geohronologiâ i Korrelâtsiâ Pozdnego Kainozoâ Paratetisa. 97 pp. Nauka, Moskva.
• Danukalova, G.A. 1990. Dvustvorčatye Mollûski iz Stratigrafiâ Akča gila. 367 pp. Ph.D. Thesis, Paleontologičeskij Institut, Akademiâ Nauk SSSR, Moskva.
• Dombrovskij, V.S. 1914. A note on the geological conditions of the location of the bones of fossil mammals in El’dar (Tiflis province) [in Russian]. Trudy Geologičeskogo Muzeâ Imeni Petra Veliokogo Imperatorskoj Akademii Nauk 8 (14): 231—238.
• Dubrovo, I.A. and Burchak-Abramovich, N.I. 1986. New data on the evolution of Bovine of the tribe Bovini. Quartärpälaontologie 6: 13–21.
• Dzigrashvili, A.T. [Dzigrašvili, A.T.] and Sepashvili, O.A. [Sepašvili, O.A.] 1978. The conditions of deposition of the Tsteltskaro limestones [in Russian]. Trudy Vsesoûznogo Neftânogo Naučno-Issledovatel’skogo Geologorazvedočnogo Instituta 115 (14): 201–209.
• Eisenmann, V. and Sondaar, P. 1989. Hipparions and the Mio-Pliocene boundary. Bulletino della Società Paleontologica Italiana 28 (2–3): 217–226.
• Eronen, J.T. and Rook, L. 2004. The Mio-Pliocene European primate fossil record: dynamics and habitat tracking. Journal of Human Evolution 47: 323–341.
• Eronen, J.T., Ataabadi, M.M., Micheels, A., Karmea, A., Bernor, R.L., and Fortelius, M. 2009. Distribution history and climatic controls of the Late Miocene Pikermian chronofauna. Proceedings of the National Academy of Sciences 106 (29): 11867–11871.
• Fataliev, R.A. 1964. Verhnesarmatskaâ Flora Gory Katar v Meždureč’e Kury i Iori. 17 pp. Synopsis of the Ph.D. Thesis, Botaničeskij institut imeni V.L. Komarova, Akademiâ Nauk SSSR, Leningrad.
• Ferring, R., Oms, O., Agustí, J., Berna, F., Nioradze, M., Shelia, T., Tappen, M., Vekua, A., Zhvania, D., and Lordkipanidze, D. 2011. Earliest human occupations at Dmanisi (Georgian Caucasus) dated to 1.85–1.78 Ma. Proceedings of the National Academy of Sciences 108 (26): 10432–10436.
• Filippova, N.Y. 1997. Palinologiâ Verhnego Pliotsena–Srednego Plejstotsena Ûga Kaspiiskoj Oblasti. 164 pp. GEOS, Moskva.
• Forte, A., Cowgill, E., Bernardin, T., Kreylos, O., and Hamann, B. 2010. Late Cenozoic deformation of Kura fold-thrust belt, southern Greater Caucasus. Geological Society of America Bulletin 122: 465–486.
• Forte, A., M., Sumner, D. Y., Cowgill, E., Stoica, M., Murtuzayev, I., Kangarli, T., Elashvili, M., Godoladze, T., and Javakhishvili, Z. 2015. Late Miocene to Pliocene stratigraphy of the Kura Basin, a subbasin of the South Caspian Basin: implications for the diachroneity of stage boundaries. Basin Research 27 (3): 247–271.
• Gabashvili, E.G. [Gabašvili, E.G.] 1956. Miocene antelopes hold in the collections of the Georgian State Museum [in Georgian with extended summary in Russian]. Vestnik Gosudarstvennogo Muzeâ Gruzii 17A: 15–26.
• Gabashvili, E.G. [Gabašvili, E.G.] 1983. Late Miocene proboscideans of Udabno [in Russian]. Vestnik Gosudarstvennogo Muzeâ Gruzii 32A: 29–56.
• Gabashvili, E.G. [Gabašvili, E.G.] 1987. History of development of the Late Miocene bovids from Udabno [in Russian]. Vestnik Gosudarstvennogo Muzeâ Gruzii 34A: 3–44.
• Gabashvili, E.G. [Gabašvili, E.G.] 2002. Gareǰis Xerxemliant’a P’auna. 86 pp. Georgian centre for prehistory researches, Tbilisi.
• Gabunia, L.K. [Gabuniâ, L.K.] 1959. K Istorii Gipparionov. 570 pp. Akademiâ Nauk SSSR, Moskva.
• Gabunia, L.K. [Gabuniâ, L.K.] and Vekua, A.K. 1963. Taribanis Namarxi Spilo. 67 pp. Akademiâ Nauk Gruzinskoj SSR, Tbilisi.
• Gabunia, L.K. and Vekua, A.K. 1998. The find of Dinopelis in the Pliocene of Georgia. Bulletin of the Georgian Academy of Sciences 157 (2): 335–338.
• Gabunia, L.K., Gabashvili, E.G., Vekua, A.K., and Lordkipanidze, D. 2001. The Late Miocene hominoid from Georgia. In: L. de Bonis, G.D. Koufos, and P.J. Andrews (eds.), Hominoid Evolution and Climate Change in Europe. Vol. 2: Phylogeny of the Neogene Hominoid Primates in Eurasia, 316–325. Cambridge University Press, Cambridge.
• Gabunia, L.K. [Gabuniâ, L.K.], Matskhonashvili, K.G. [Matshonašvili, K.G.], and Chkheidze, D.V. [Čheidze, D.V.] 1966. On the age of continental deposits of the Iagluja Mountain [in Georgian, with abstract in Russian]. Soobŝeniâ Akademii Nauk Gruzinskoj SSR 43 (1): 129–132.
• Gadzhiev, D.V. [Gadžiev, D.V.] 1996. El’darskaâ Verhnesarmatskaâ Gipparionovaâ Fauna. 344 pp. ELM, Baku.
• Gagnidze, R. and Davitadze, M. 2000. Adgilobrivi P’lora (Sak’art’velos Mc’enaret’a Samqaro). 274 pp. Ač’ara, Batumi.
• Gamkrelidze, I., Gamkrelidze, M., Loladze, M., and Tsamalashvili, T. 2015. New tectonic map of Georgia (explanatory note). Bulletin of the Georgian National Academy of Sciences 9 (1): 211–216.
• Ganzej, S.S. 1987. Late Caenosoic deposits of Ponto-Caspian and fission-treck dating of ash layers [in Russian]. In: A.M. Korotkij (ed.), Metod Trekov v Geologii i Geografii, 34–45. Dal’nevostočnyj Naučnyj Centr, Tihookeanskij Institut Geografii, Akademiâ Nauk SSSR, Vladivostok.
• Golubjatnikov, D.V. [Golubâtnikov, D.V. ] 1927. Detailed geological map of Absheron peninsula. Atashan district [in Russian]. Trudy Geologičeskoj Komissii, Novaâ Seriâ 130: 1–314.
• Grichuk, V.P. [Gričuk, V.P.] 1969. Glacial floras and their classification [in Russian]. In: I.P. Gerasimov (ed.), Poslednij Lednikovyj Pokrov Na Severo-Zapade Evropeiskoj Časti SSSR, k VIII Kongressu INQUA Pariž 1969, 57–70. Nauka, Moskva.
• Gromov, V.I. 1948. Palaeontological and archaeological background for the stratigraphy of the continental Quaternary deposits on USSR territory [in Russian]. Trudy Instituta Geologičeskih Nauk AN SSSR, Seriâ Geologičeskaâ 64 (17): 1–521.
• Grossheim, A.A. 1948. Rastitel’nyj Pokrov Kavkaza. 267 pp. Akademiâ Nauk SSSR, Moskva.
• Gurarij, G.Z., Kudasheva, I.A., Trubikhin, V.M., Fein, A.G., and Shipunov, S.V. 1995. Age of the Natskhor and Shiraki Formations of Eastern and Central Georgia and the problem of the Miocene–Pliocene boundary—paleomagnetism and stratigraphy. Stratigraphy and Geological Correlation 3: 156–166.
• Hemmer, H., Kahlke, R.-D., and Vekua, A.K. 2004. The Old World puma – Puma pardoides (Owen, 1846) (Carnivora: Felidae)—in the lower Villafranchian (upper Pliocene) of Kvabebi (East Georgia, Transcaucasia) and its evolutionary and biogeographical significance. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 233: 197–231.
• Hilgen, F., Lourens, L., Van Dam, J.A., Beu, A., Boyes, A.F., Cooper, R., Krijgsman, W., Ogg, J., Piller, W., and Wilson, D. 2012. The Neogene Period. In: F.M. Gradstein, J.G. Ogg, M.D. Schmitz, and G.M. Ogg (eds.), The Geologic Time Scale, 923–978. Elsevier, Boston.
• Isaeva, M.I. and Mamedova, N.R. 1989. Paleomagnetic studies of the major sections of the Middle Kura depression (Pliocene–Pleistocene) [in Russian]. Izvestiâ Akademii Nauk Azerbajdžanskoj SSR, Seriâ Nauka o Zemle 3: 123–127.
• Khuchua, M.F. [Hučua, M.F.] and Shatirishvili, T.M. [Šatirišvili, T.M.] 1937. Mineralogical composition of sandy deposits of the upper Sarmatian and Shiraki Formation in the sections of Akhtatapa, Kotsakhuriskedi [in Russian]. Azerbajdžanskoje Neftânoe Hozâistvo 2: 9–17.
• Koiava, K., Maissuradze, L., Strasser, A., Shatilova, I., Kvaliashvili, L., and Glonti, V. 2012. Palaeogeography of the Sarmatian of eastern Georgia. Bulletin of the Georgian National Academy of Sciences 6 (3): 91–98.
• Kolesnikov, V.P. 1940. Scheme of stratigraphic subdivision of Neogene deposits of the Ponto-Caspian region [in Russian]. In: A.D. Arhangel’skij (ed.), Neogen. Stratigrafiâ SSSR 12: 567–574.
• Kolesnikov, V.P. 1950. Akčagil’skie i Apšeronskie Mollûski. Paleontologiâ SSSR, Vol. 10, Part 3 (12). 259 pp. Akademiâ Nauk SSSR, Moskva.
• Kostopoulos, D.S. 2014. Taxonomic reassessment and phylogenetic relationships of Miocene homonymously spiral-horned antelopes. Acta Palaeontologica Polonica 59 (1): 9–29.
• Kostopoulos, D.S. and Bernor, R.L. 2011. The Maragheh bovids (Mammalia, Artiodactyla): systematic revision and biostratigraphiczoogeographic interpretation. Geodiversitas 33 (4): 649–708.
• Kudrjavtsev, N.A. [Kudrâvtsev, N.A.] 1938. Geological research in the Alazani and Kura region (brief preliminary report for the years 1930–1931) [in Russian]. Trudy Neftânogo Geologo-Razvedovočnogo Instituta Seriâ B 32:1–58.
• Kuzmina, I.E. and Sablin, M.V. 1991. On new finds of fossil remains of Equus stenonis Cocchi in the South Caucasus [in Russian]. Trudy Zoologičeskogo Instituta Akademii Nauk SSSR 238: 61–67.
• Lebedeva, N.A. 1972. Geological position of the remains of terrestrial mammals of Khapry, Taman’ and Tiraspol faunistical complexes in the cut of marine layers of Akchagyl and Apsheron of eastern Transcaucasia [in Russian]. Byulleten’ Komissii po Izučeniû Četvertičnogo Perioda 38: 99–117.
• Lebedeva, N.A. 1974. Korrelâtsiâ Morskih i Kontinental’nyh Otloženij Eopleistotsena i Nižnego Pleistotsena Ponto-Kaspijskoj Oblasti. 53 pp. Synopsis of the Ph.D. Thesis, Geologičeskij institut, Akademiâ Nauk SSSR, Moskva.
• Lebedeva, N.A. 1978. Korrelâciâ Antropogenovyh Tolŝ Ponto-Kaspiâ. 135 pp. Nauka, Moskva.
• Leroy, S.A.G., Tudryn, A., Chalié, F., and López-Merino, L. 2013. From the Allerød to the mid-Holocene: palynological evidence from the South Basin of the Caspian Sea. Quaternary Science Reviews 78: 77–97.
• Lordkipanidze, D., Jashashvili, T., Vekua, A., Ponce de León, M.S., Zollikofer, C.P.E., Rightmire, G.P., Pontzer, H., Ferring, R., Oms, O., Tappen, M., Bukhsianidze, M., Agusti, J., Kahlke, R.-D., Kiladze, G., Martinez-Navarro, B., Mouskhelishvili, A., Nioradze, M., and Rook, L. 2007. Postcranial evidence from early Homo from Dmanisi, Georgia. Nature 449: 305–310.
• Maisadze, F. 2013. Duration of Cenozoic orogenies (on the example of Georgia). Bulletin of the Georgian National Academy of Sciences 7 (2): 97–104.
• Meladze, G.K. 1967. Gipparionovaâ Fauna Arkneti i Bazaleti. 168 pp. Mecsniereba, Tbilisi.
• Meladze, G.K. 1985. Obzor Gipparionovyh Faun Kavkaza. 75 pp. Mecniereba, Tbilisi.
• Mauvilly, J., Koiava, K., Gamkrelidze, I., and Mosar, J. 2016. Tectonics in the Georgian Greater Caucasus: a structural cross-section in an inverted rifted basin setting. In: S. Castelltort, S. Girardclos, E. Gnos, E. Samankassou, S. Scmits, M. Lupi, C. Frischkencht, M. Chiradia, G. Simpson , R. Martini, U. Schaltegger, and D. Ariztegui (eds.), 14th Swiss Geoscience Meeting, Geneva, Switzerland, November 18–19, 33–34. Swiss Academy of Sciences, Geneva. [published online, doi: 10.13140/RG.2.2.26540.56963]
• Mauvilly, J. 2017. Tectonics in the Georgian Greater Caucasus: a Structural Cross Section in an Inverted Rifted Basin Setting. 127 pp. Unpublished M.Sc. Thesis, University of Fribourg, Fribourg.
• Nanda, A.C. 2002. Upper Siwalik mammalian faunas of India and associated events. Journal of Asian Earth Sciences 21: 47–58.
• Nargolwalla, M.C. 2009. Eurasian Middle and Late Miocene Hominoid Paleobiogeography and the Geographic Origins of the Homininae. 534 pp. Ph.D. Thesis, University of Toronto Library and Archives Canada, Toronto.
• Nemčok, M., Glonti, B., Yukler, A., and Marton, B. 2013. Development history of the foreland plate trapped between two converging orodens; Kura valley, Georgia, case study. Geological Society, London, Special Publications 377: 159–188.
• Nevesskaya, L.A. [Nevesskaâ, L.A.] and Trubikhin, V.M. [Trubihin, V.M.] 1984. The history of the Caspian Basin and its fauna of mollusks in the Late Pliocene and Early Pleistocene [in Russian]. In: V.L. Jahimovič (ed.), Antropogen Evrazij, 19–27. Nauka, Moskva.
• Nikiforova, K.V., Krasnov, I.I., Aleksandrova, L.P., Vasiliev, Y.M., Konstantinova, N.A., and Tshepalyga, A.L. [Tšepalyga, A.L.] 1982. Chronostratigraphic scheme of the Late Pliocene–Antropogene deposits of the European part of the USSR [in Russian]. In: D.V. Nalivkin and B.S. Sokolov (eds.), Stratigrafiâ SSSR, Četvertičnaâ Sistema 1, unnumbered insert. Nedra, Moskva.
• Nomade, S., Pastre, J.F., Guillou, H., Faure, M., Guérin, C., Delson, E., Debard, E., Voinchet, P., and Messager, E. 2014. 40Ar/39Ar constraints on some French landmark Late Pliocene to Early Pleistocene large mammalian paleofaunas: Paleoenvironmental and paleoecological implications. Quaternary Geochronology 21: 2–15.
• Ogg, J.G. 2012. Geomagnetic Polarity Time Scale. In: F.M. Gradstein, J.G. Ogg, M.D. Schmitz, and G.M. Ogg (eds.), The Geologic Time Scale, 85–114. Elsevier, Boston.
• Orlov, Y.A. 1941. A new locality of hipparion-fauna at Djaparidze, east Georgia [in Russian with abstract in English]. Trudy Paleontologičeskogo Instituta Akademii Nauk SSSR 8 (3): 99–101.
• Palmqvist, P., Martínez-Navarro, M., Pérez-Claros, J.A., Torregrosa, V., Figueirido, B., Jiménez-Arenas, J.M., Espigares, M.-P., Ros-Montoya, S., De Renzi, M. 2011. The giant hyena Pachycrocuta brevirostris: Modelling the bone-cracking behavior of an extinct carnivore. Quaternary International 243 (1): 61–79.
• Pevzner, M.A. 1982. Application of paleomagnetic method for the stratigraphy of Quaternary deposits [in Russian]. In: D.V. Nalivkin and B.S. Sokolov (eds.), Stratigrafiâ SSSR, Četvertičnaâ Sistema 1, 149–154. Nedra, Moskva.
• Pickford, M. 2013. Re-assessment of the suids from the Sables marins de Montpellier and selection of a lectotype for Sus provincialis Blainville, 1847. Geodiversitas 35 (3): 655–689.
• Popov, G.I. 1969. Stratigraphic scheme of the Akchagylian stage [in Russian]. Geologiâ i Poleznie Iskopaemye Turkmenii 2: 179–224.
• Radionova, E.P., Golovina, L.A., Filippova N.Y., Trubikhin, V.M., Popov, S.V., Goncharova, I.A., Vernigorova, Y.V., and Pinchuk, T.N. 2012. Middle–Upper Miocene stratigraphy of the Taman Peninsula, Eastern Paratethys. Central European Journal of Geosciences 4 (1): 188–204.
• Riabinin, A. 1913. Recherches géologiques dans la region de la steppe de Chiraki et de ses alentours [in Russian with summary in French]. Trudy Geologičeskogo Komiteta, Novaâ Seriâ 93: 1–73.
• Rook, L., Bartolini Lucenti, S., Bukhsianidze, M., and Lordkipanidze, D. 2017. The Kvabebi Canidae record revisited (Late Pliocene, Sighnaghi, eastern Georgia). Journal of Paleontology 91 (6): 1258–1271.
• Ruzhentsev, V.E. [Ružentsev, V.E.] 1928. The division of the Apsheronian stage based on the study of the Apsheronian deposits of the Lesser Kharami Range [in Russian]. Bûlleten’ Moskovskogo Obŝestva Ispytatelej Prirody, Otdel Geologičeskij, Novaâ Seriâ 36 (6): 241–266.
• Sablin, M.V. 1990. Remains of carnivores and ungulates from the Lower Absheronian deposits of Azerbaidjan [in Russian]. Trudy Zoologičeskogo Instituta Akademii Nauk SSSR 213: 138–142.
• Safarov, I.S. 1979. Subtropičeskie Lesa Talyša. 157 pp. ELM, Baku.
• Sahni, M.R. and Khan, E. 1968. Probison dehmi n. g. n. sp. a recent find of an upper Sivalik bovid. Bayerische Staatssammlung für Paläontologie und Historische Geologie 8: 247–251.
• Semenov, Y.A. 1989. Iktiterii i Morfologičeski Shodnye Geny Neogena SSSR. 178 pp. Naukova Dumka, Kiev.
• Sen, S. 1997. Magnetostratigraphic calibration of the European Neogene mammal chronology. Palaeogeography, Palaeoclimatology, Palaeoecology 133: 181–204.
• Shatilova, I. and Kokolashvili, I. 2011. The flora and vegetation of eastern Georgia in the Sarmatian. Proceedings of the Georgian Academy of Sciences, Biological Series B 9 (1–4): 65–70.
• Shatilova, I. and Kokolashvili, I. 2012. The comparison of eastern and western Georgia’s Sarmatian vegetation and climate. Proceedings of the Georgian National Museum, Natural Sciances and Prehistory Section 4: 31–36.
• Shatilova, I. [Šatilova, I.] and Kokolashvili, I. [Kokolašvili, I.] 2013. Palinologičeskaâ Harakteristika Sarmatskih Otloženii Vostočnoj Gruzii [in Russian with English summary]. 110 pp. Universal, Tbilisi.
• Shatilova, I., Maissuradze, L., Kokolashvili, I., and Koiava, K. 2012. The comparison of the results of palynological and microfaunistical investigations of the Sarmatian deposits of eastern Georgia. Bulletin of the Georgian National Academy of Sciences 6 (2): 139–144.
• Shatilova, I., Mchedlishvili, N., Rukhadze, L., and Kvavadze, E. 2011. The History of the Flora and Vegetation of Georgia (South Caucasus). 200 pp. Universal, Tbilisi.
• Shchelinsky, V.E. [Ŝelinskij, V.E.] 2014. Eopleistotsenovaâ Rannepaleolitičeskaâ Stoânka Rodniki 1 v Zapodnom Predkavkaz’e. 168 pp. IIMC RAS, Saint-Petersburg.
• Shchelinsky, V.E., Gurova, M., Tesakov, A.S., Titov, V.V., Frolov, P.D., and Simakova, A.N. 2016. The Early Pleistocene site of Kermek in western Ciscaucasia (southern Russia): Stratigraphy, biotic record and lithic industry (preliminary results). Quaternary International 393: 51–69.
• Sotnikova, M.V. 1989. Hiŝnye Mlekopitaûŝie Pliotsena—Rannego Plejstotsena. 123 pp. Nauka, Moskva.
• Sotnikova, M.V. and Rook, L. 2010. Dispersal of the Canini (Mammalia, Canidae: Caninae) across Eurasia during the Late Miocene to Early Pleistocene. Quaternary International 212: 86–97.
• Sotnikova, M.V. and Sablin, M.V. 1993. The late Villafranchian association of carnivorous mammals from the locality Palan-Tyukan [in Russian with abstract in English]. Trudy Zoologičeskogo Instituta Aka demii Nauk SSSR 249: 134–145.
• Sultanov, K.M. 1964. Apšeronskij Ârus Azerbajdžana. 233 pp. Azernašr, Baku.
• Tappen, M., Adler, D.S., Ferring, C.R., Gabunia, M., Vekua, A., and Swisher III, C.C. 2002. Akhalkalaki: The taphonomy of an Early Pleistocene locality in the Republic of Georgia. Journal of Archaeological Science 29: 1367–1391.
• Tarkhnishvili, D. 2014. Historical Biogeography of the Caucasus. 180 pp. Nova Science Publishers Inc., New York.
• Tesakov, A.S. 2004. Biostratigraphy of middle Pliocene–Eopleistocene of Eastern Europe (based on small mammals) [in Russian]. Trudy Geologičeskogo Instituta 554: 1–247.
• Tesakov, A.S., Titov, V.V., Simakova, A.N., Florov, P.D., Syromyatnikova, E.V., Kurshakov, S.V., Volkova, N.V., Trikhunkov, Y.I., Sotnikova, M.V., Kruskop, S.V., Zelenkov, N.V., Tesakova, E.M., and Palatov, D.M. 2017. Late Miocene (Early Turolian) vertebrate faunas and associated biotic record of the Northern Caucasus: geology, taxonomy, palaeoenvironment, biochronology. Fossil Imprint 73 (3–4): 383–444.
• Trubikhin, V.M. [Trubihin, V.M.] 1977. Paleomagnetizm i stratigrafiâ akčagil’skih otloženii Zapodnoj Turkmenii. 77 pp. Nauka, Moskva.
• Tsiskarishvili, G.V. [Tsiskarišvili, G.V.] 1987. Pozdnetretičnye Nosorogi (Rhinocerotidae) Kavkaza. 142 pp. Mecniereba, Tbilisi.
• Utescher, T., Erdei, B., Hably, L., and Mosbrugger, V. 2017. Late Miocene vegetation of the Pannonian Basin. Palaeogeography, Palaeoclimatology, Palaeoecology 467: 131–148.
• Van Baak, C.G.C., Vasiliev, I., Stoica, M., Kuiper, K.F., Forte A.M., Aliyeva, E., and Krijgsman, W. 2013. A magnetostratigraphic time frame for Plio-Pleistocene transgressions in the South Caspian Basin, Azerbaijan. Global and Planetary Change 103: 119–134.
• Vangengeim, E.A. and Tesakov, A.S. 2008. Late Sarmatian mammal localities of the Eastern Paratethys: Stratigraphic position, magnetochronology, and correlation with the European continental scale. Stratigraphy and Geological Correlation 16 (1): 92–103.
• Vangengeim, E.A., Muskhelishvili, L.V. [Mušelišvili, L.V.], Pevzner, M.A., and Tsiskarishvili, G.V. [Tsiskarišvili, G.V.] 1988. Place of the upper boundary of Vallesian on the stratigraphic scheme of the Eastern Paratethys [in Russian]. Izvestiâ Akademii Nauk SSSR, Seriâ Geologičeskaâ 6: 67–74.
• Vangengeim, E.A., Gabunia, L.K. [Gabuniâ, L.K.], Pevzner, M.A., and Tsiskarishvili, G.V. [Tsiskarišvili, G.V.] 1989. Stratigraphic position of the sites with hipparion faunas from the South Caucasus in the light of the data of magnetostratigraphy [in Russian]. Izvestiâ Akademii Nauk SSSR, Seriâ Geologičeskaâ 8: 70–77.
• Vanishvili, N., Vekua, A.K., and Kiladze, G. 2007. Simocyon from the Late Miocene of Georgia. Problems of Paleobiology 2: 142–146.
• Vasiliev, I., Iosifidi, A.G., Khramov, A.N., Krijgsman, W., Kuiper, K., Langereis, C.G., Popov, V.V., Stoica, M., Tomsha, V.A., and Yudin, S.V. 2011. Magnetostratigraphy and radio-isotope dating of Upper Miocene–Lower Pliocene sedimentary successions of the Black Sea Basin (Taman Peninsula, Russia). Palaeogeography, Palaeoclimatology, Palaeo ecology 310 (3): 163–175.
• Vekua, A.K. 1972. Kvabebskaâ Fauna Akčagil’skih Pozvonočnyh. 351 pp. Nauka, Moskva.
• Vekua, A.K. 1991. History of the animal world—vertebrates [in Russian]. In: L.I. Maruašvili (ed.), Gruziâ v Antropogene, 308–339. Sak’art’velo, Tbilisi.
• Vekua, A.K. and Lordkipanidze, D. 1998. The Pleistocene paleoenvironment of the Transcaucasus. Quaternaire 9 (4): 261–266.
• Vekua, A.K., Lordkipanidze, D., and Bukhsianidze, M. 2009. Importance of Diliska Vertebrate Fauna for the Biostratigraphy of South Caucasus. Bulletin of the Georgian National Academy of Sciences 3 (2): 130–135.
• Vekua, A.K., Lordkipanidze, D., Bukhsianidze, M., and Chagelishvili, R. 2010. Kotsakhuri (eastern Georgia) Pliocene fauna of vertebrates and Taribana elephant. Proceedings of the Georgian Academy of Sciences, Biological Series B 8 (1–2): 79–84.
• Vekua, A.K. and Trubikhin, V.M. [Trubihin, V.M.] 1988. New site of fossil mammals in eastern Georgia [in Russian with Georgian and English abstract]. Soobŝeniâ Akademii Nauk Gruzinskoj SSR 132 (1): 197–200.
• Vislobokova, I.A. 2008. The major stages in the evolution of artiodactyl communities from the Pliocene–early Middle Pleistocene of northern Eurasia: Part 1. Paleontological Journal 42: 297–312.
• Vislobokova, I.A. and Tesakov, A.S. 2013. Early and Middle Pleistocene of Northern Eurasia. In: S.A. Elias (ed.), The Encyclopedia of Quaternary Sciences, 605–614. Elsevier, Amsterdam.
• Werdelin, L. 2001. A revision of the genus Dinofelis (Mammalia, Felidae). Zoological Journal of the Linnean Society 132: 147–258.
• Zohary, M. 1973. Geobotanical Foundations of the Middle East. 739 pp. Verlag, Stuttgart.

Identyfikatory

DOI

10.4202/app.00499.201