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The lacustrine oil shales of the Coal Creek Member of the Kishenehn Formation in northwestern Montana comprise a relatively unstudied middle Eocene fossil insect locality. Herein, we detail the stratigraphic position of the fossiliferous unit, describe the insect fauna of the Coal Creek locality and document its bias towards very small but remarkably preserved insects. In addition, the depositional environment is examined and the mineral constituents of the laminations that comprise the varves of the Kishenehn oil shale are defined. Fifteen orders of insects have been recorded with the majority of all insects identified as aquatic with the families Chironomidae (Diptera) and Corixidae (Hemiptera) dominant. The presence of small aquatic insects, many of which are immature, the intact nature of >90% of the fossil insects and the presence of Daphnia ephippia, all indicate that the depositional environment was the shallow margin of a large freshwater lake. The fossil insects occur within fossilized microbial mat layers that comprise the bedding planes of the oil shale. Unlike the fossiliferous shales of the Florissant and Okanagan Highlands, the mats are not a product of diatomaceous algae nor are diatom frustules a component of the sediments or the varve structure. Instead, the varves are composed of very fine eolian siliciclastic silt grains overlaid with non-diatomaceous, possibly cyanobacteria-derived microbial mats which contain distinct traces of polyaromatic hydrocarbons. A distinct third layer composed of essentially pure calcite is present in the shale of some exposures and is presumably derived from the seasonal warming-induced precipitation of carbonate from the lake's waters. The Coal Creek locality presents a unique opportunity to study both very small middle Eocene insects not often preserved as compression fossils in most Konservat-Lagerstätte and the processes that led to their preservation.
A unique dinosaur assemblage from the Cretaceous beds of western Inner Mongolia preserves geologic and paleontologic data that clearly delineate both the timing and mechanism of death. Over twenty individuals of the ornithomimid Sinornithomimus dongi perished while trapped in the mud of a drying lake or pond, the proximity and alignment of the mired skeletons indicating a catastrophic mass mortality of a social group. Histologic examination reveals the group to consist entirely of immature individuals between one and seven years of age, with no hatchlings or mature individuals. The Sinornithomimus locality supports the interpretation of other, more taphonomically ambiguous assemblages of immature dinosaurs as reflective of juvenile sociality. Adults of various nonavian dinosaurs are known to have engaged in prolonged nesting and post hatching parental care, a life history strategy that implies juveniles spent considerable time away from reproductively active adults. Herding of juveniles, here documented in a Cretaceous ornithomimid, may have been a common life history strategy among nonavian dinosaurs reflecting their oviparity, extensive parental care, and multi−year maturation.
New material collected from the Kliphoek Member of the Nama Group (Kuibis Subgroup, Dabis Formation) on Farm Aar, southern Namibia, offers insights concerning the morphology of the Ediacaran organism Pteridinium. Pteridinium fossils previously described as being preserved in situ have been discovered in association with scour−and−fill structures indicative of transport. Additionally, two Pteridinium fossils have been found within sedimentary dish structures in the Kliphoek Member. A form of organic surface with a discrete membrane−like habit has also been recovered from Farm Aar, and specimens exist with both Pteridinium and membrane−like structures superimposed. The association between Pteridinium fossils and membrane−like structures suggests several possibilities. Pteridinium individuals may have been transported before burial along with fragments of microbial mat; alternately they may have been enclosed by an external membranous structure during life.
From the Upper Miocene (Messinian) of the Alba area in Piedmont, northern Italy, are fossil dragonflies (mainly larvae) described, which were preserved in transparent gypsum crystals. The specimens belong probably to a single species (Oryctodiplax gypsorum), but occur in various pre-imaginal stages. This individual-rich but species-poor palaeoentomofauna appears to be almost autochthonous in origin and may have developed under hypersalinar conditions in a lagoonal environment during the Messinian salinar event.
Amphiesmenopterans are represented in the Lower Toarcian marine insect taphocoenoses of Germany by stem group members (Necrotaulius HANDLIRSCH, 1906, Mesotrichopteridium HANDLIRSCH, 1906). Female terminalia of Necrotaulius resembling the ovipositor indicate that these insects laid their eggs rather in soil than in water. One species of genuine Trichoptera (Liadotaulius HANDLIRSCH, 1939) exhibited sexual dimorphism. At least two species of Lepidoptera related to Micropterygidae were found.
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Upper Cretaceous amniotic eggs from Gobi Desert

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There are described several types of Upper Cretaceous fossil amniote eggs, clutches and eggshells, collected during the Polish-Mongolian Paleontological Expeditions (1963—1971) in several localities of the Gobi Desert. Isotopic ratios of C and О are reported and discussed. The eggs may be attributed to saurischian and ornithischian dinosaurs, and palaeognathous birds. Four types of problematica are also described. Taphonomy and restoration of eggshells’ diffusive properties as compared with modern hard-shelled eggs suggest paleoecological interpretation of the nesting conditions as rather dry for smooth-shelled eggs, humid for ornamented eggs, and very humid for the thickest-shelled eggs. Different nest types are postulated for ornamented and smooth eggs, based on taphonomy and shell parameters.
Diverse carbonaceous microfossils, including exceptionally preserved remains of non−biomineralizing metazoans, are reported from a basal middle Cambrian interval of the Kaili Formation (Guizhou Province, China). The application of a gentle acid maceration technique complements previous palynological studies by revealing a larger size−class of acritarchs, a richer assemblage of filamentous microfossils, and a variety of previously unrecovered forms. Metazoan fossils include Wiwaxia sclerites and elements derived from biomineralizing taxa, including chancelloriids, brachiopods and hyolithids, in common with previously studied assemblages from the early and middle Cambrian of Canada. In addition, the Kaili Formation has yielded pterobranch remains and an assemblage of cuticle fragments representing “soft−bodied” worms, including a priapulid−like scalidophoran. Our results demonstrate the wide distribution and palaeobiological importance of microscopic “Burgess Shale−type” fossils, and provide insights into the limitations and potential of this largely untapped preservational mode.
Latest Emsian (Early Devonian) sediments at the famous mud−moundand trilobite−locality Hamar Laghdad (Tafilalt, Morocco) yielded some red−coloured remains of phacopid trilobites. Closer examination revealed that the eyes of these phacopids are often greenish in colour. EDX−analyses showed that the lenses retained their original calcitic composition, possibly greenish due to Feand Mn−impurities, while most of the exoskeleton was silicified. The silicified parts contain elevated concentrations of iron which causes the red colour. This phenomenon is explained by the porosity of the exoskeleton in contrast to the homogeneous and massive construction of the lenses and their Mg−content. These incompletely silicified trilobites enabled a reconstruction of the silicification process in trilobites. Their diagenetic alteration probably occurred as a result of events associated with the Cretaceous transgression.
Fossil catshark egg capsules, Scyliorhinotheca goederti gen. et sp. nov., are reported from a Late Eocene deep−water methane−seep calcareous deposit in western Washington State, USA. The capsules are preserved three−dimensionally and some show mineralized remnants of the ribbed capsule wall consisting of small globular crystals that are embedded in a microsparitic matrix. The globules are calcitic, but a strontium content of 2400–3000 ppm suggests that they were origi− nally aragonitic. The carbonate enclosing the egg capsules, and the capsule wall itself, show 13C values as low as −36.5‰, suggesting that formation was induced by the anaerobic oxidation of methane and hence in an anoxic environ− ment. We put forward the following scenario for the mineralization of the capsule wall: (i) the collagenous capsules expe− rienced a sudden change from oxic to anoxic conditions favouring an increase of alkalinity; (ii) this led to the precipitation of aragonitic globules within the collagenous capsule wall; (iii) subsequently the remaining capsule wall was mineralized by calcite or aragonite; (iv) finally the aragonitic parts of the wall recrystallized to calcite. The unusual globular habit of the early carbonate precipitates apparently represents a taphonomic feature, resulting from mineralization mediated by an organic matrix. Taphonomic processes, however, are at best contributed to an increase of alkalinity, which was mostly driven by methane oxidation at the ancient seep site.
Five cephalopod specimens from the Lower Devonian of Bohemia (Czech Republic) preserve colour patterns. They include two taxonomically undeterminable orthoceratoids and three oncocerid nautiloids assigned to the genus Ptenoceras. The two fragments of orthocone cephalopods from the lowest Devonian strata (Lochkovian, Monograptus uniformis Zone) display colour patterns unusual in orthoceratoids. They have irregular undulating and zigzag strips that are preserved on counterparts of adapertural regions of specimens flattened in shale, despite their original aragonitic shell having been completely dissolved. These are probably the result of the proteinous pigment inside the shell wall, being substituted during diagenesis by secondary minerals leaving only an altered trace of the original shell. Orthoceratoids from sediments unsuitable for preservation of this feature discussed here thus demonstrate an exceptional case of preservation of colour patterns, not only within Devonian cephalopods but also within other Devonian molluscs. Three specimens of Ptenoceras that preserve colour patterns come from younger Lower Devonian strata. Oblique spiral adaperturally bifurcating bands are preserved in P. alatum from the Pragian and zigzags in P. nudum from the Dalejan. Juvenile specimen of Ptenoceras? sp. from the Pragian exhibits highly undulating transversal bands—a pattern resembling colour markings in some Silurian oncocerids. Dark grey wavy lines observed on the superficially abraded adapical part of a phragmocone of nautiloid Pseudorutoceras bolli and interpreted formerly to be colour markings are here reinterpreted as secondary pigmented growth lines. Other Devonian fossils including a single brachiopod and several gastropods from the Barrandian Area with preserved colour patterns are mentioned. Variety of cephalopod colour patterns, their taxonomic significance, function and significance for palaeoecological interpretation, palaeoenvironmental conditions favouring colour pattern preservation and systematic affiliation of taxa with colour pattern preserved are discussed.
Considering the fossil record, foraminifers constitute one of the most ubiquitous groups in marine deposits due to a high preservation potential of tests. Moreover, borings appear to be widely distributed in the tests. Findings of the borings have potential usefulness as palaeoecological markers and can be significant for taphonomical studies of foraminifers. Formal naming of the borings is needed to further their usefulness, and therefore a new ichnotaxon Curvichnus semorbis igen. et isp. nov. is erected.
A new preservation of aptychi is described from the Valanginian limestone−marl alternations of the Vergol section (Drôme), located in the Vocontian Basin (SE France). Aptychi are arranged into two parallel rows which are generally 50 mm in length and separated by 4 mm. The alignments are very often made by entire aptychi (around 10 mm in length), oriented following their harmonic margin. Aptychi show the outside of valve to the viewer: they are convex−up. This fossilization of aptychi is successively interpreted as the result of post−mortem transport by bottom currents (taphonomicresedimentation process) or the residues (bromalites: fossilized regurgitation, gastric and intestinal contents, excrement) from the digestive tract of an ammonoid−eater (biological processes). Both the parallel rows of aptychi are more likely interpreted as a coprolite (fossil faeces) and they could be considered as both halves (hemi−cylindrical in shape) of the same cylindrical coprolite which would have been separated in two parts (following the long axis) just after the animal defecated. Considering this hypothesis, a discussion is proposed on the hypothetical ammonoid−eater responsible for them.
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