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The objective of the study was to compare the seven tree species in terms of their role in the formation of a complex vertical structure in mixed broadleaved stands. The data came from 18 multi−storey old−growth forests representing different communities of lowland deciduous forests in the Niepołomice Forest. In each stand, squared sample plot (80×80 m) was established. The height of trees of dbh≥7 cm was measured. Then, for each studied species, values of the two structural indices were determined. The structural diversity index (ZS) quantifies tree height variation within individual population, whereas the index of structure−forming role (RS) expresses the contribution of the given population into vertical structural diversity of the whole stand. The greatest tendency to form a population with a large height diversity belonged to Tilia cordata, Fraxinus excelsior and Ulmus laevis. Significant opportunities in this regard, however, were exhibited by other tree species, including shade−intolerant Pinus sylvestris and Alnus glutinosa. This phenomenon in the conditions of eutrophic habitats, with the constantly present competitive pressure from shade−tolerant species, should be considered as an exceptional. A major contribution to vertical structural diversity of the stand was often an attribute of the populations with a low internal height diversity, which concerned especially Quercus robur. This means that an important ecological mechanism responsible for the formation of a multi−layer canopy is complementary filling the space above ground by trees of the species with different light requirements. However, the lack of correlation between RS and ZS indices suggests that the ability of particular species to form the population with a high internal diversity is equally important. The results prove the possibility of forming stands with a very complex vertical structure in the conditions of lowland mixed deciduous forests. However, in terms of sustainable maintenance of such stand structure, the issue of the growth rate of trees at different competitive pressure and the nature of their response to the release at different stages of ontogeny remains open.
The paper describes changes, which took place over the last 30 years in old−growth forest stands occupying two permanent research plots, established in the Kaliszki and Sieraków strict protection areas in the Kampinos National Park (central Poland). Both plots have a form of ecological transects. The Kaliszki plot is 20 m wide and 700 m long (1.40 ha), while the Sieraków plot (total area of 2.56 ha) consists of two parts: main (40×460 m) and side (40×180 m) transect. In the Kaliszki plot, the stand measurements were conducted in 1993, 2007, and 2017, while in the Sieraków plot – in 1989, 1994, 2006, and 2017. For every tree with breast height diameter (DBH) ≥5 cm, species identity, DBH and spatial coordinates were determined. Starting from the second census, all trees which had died (‘losses’), as well as trees which exceeded the DBH threshold (‘gains’) since the previous record were noted, as well. The major tree species occurring on sample plots are Scots pine (Pinus sylvestris), pedunculate oak (Quercus robur), silver and downy birch (Betula sp.), and black alder (Alnus glutinosa). During the study period, the overall tree density declined by ca. 50%. The diminishing trends occurred for all dominant tree species. In contrast to major, dominant tree species, the demographic status of a group of minor tree species (lime, hornbeam and maple) was much more stable. One may even speak about some expansion in this case. Particularly, hornbeam density increased rapidly in the period 2006−2017 in some areas of the main transect located in the Sieraków plot. The future studies will show if this tendency will be maintained and will include other regions of sample plots. Assuming a lack of significant disturbances, one may expect that the stands growing on research plots will rather slowly change in the nearest future. Most dynamics will have a quantitative and not qualitative character. One may anticipate, for example, a growing role of pedunculate oak at the expense of Scots pine, however, considering that the two species are rather long−living, they will probably for a long time keep their dominant roles in the stand canopies.
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