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This paper focuses on the issue related to the response of beech leaves to the opening of stand canopy resulting from a shelterwood cut of various intensity. Four research plots each with a different stand density provided different conditions for the development of foliage. The research was conducted on dominant, codominant and subdominant sample trees representing the mean stand diameter of all plots. The data were compared for the upper, middle and lower layer of tree crowns (each equals to one third of crown length). As for dominant and codominant trees, mean leaf area showed a tendency to increase towards the lower crown layer. As for subdominant trees, relation between the location of leaves in the crown and leaf area was similar to dominant and codominant trees but this trend was observed only on the plot most open. Only in the case of codominant trees the leaf area was decreasing with a decreasing stand density. Greater stand opening results in a proportionally smaller leaf area. Growing conditions for subdominant tree leaves were completely different when compared to conditions for upper tree classes. On the plots with harvest cutting we observed that a decreasing stand density resulted in a general increase of leaf size. The only exception to the rule was the control plot where the parameters recorded were higher and inconsistent with the above described trend.
We studied the response in growth and phenology of naturally regenerated beech seedlings to changed ecological conditions over 14 and 19-years after cutting with different intensity. Five different types of stand densities were modelled: plot C – control plot – no cut, L – low intensity cut, M – medium intensity cut, H – high intensity cut and CC – clear cut, with 1, 8, 22, 53 and 100% of relative irradiation, and 66, 68, 78, 92 and 100% of through fall, respectively. We were focussing on tree height growth and leaf area. Our phenological observations were aimed at onset and course of two spring vegetative phenophases: bud-burst and leaf unfolding. Already in two-year-old beech seedlings we found significant differences in height growth; the differences in mean leaf area, however, were observed later. From the viewpoint of phenotypic plasticity, the height growth in beech seedlings represented more sensitive response to the environment than the leaf area. According to leaf area size and height growth in the beech seedlings on control plot, the stress conditions were indicated, primarily from the lacking light. With stand opening, the development of recruitment was getting better, and beginning with plot M the increase of seedlings height and leaf area became continuously related to the amount of radiation. The results of phenological observations showed that the spring phenophases in the seedlings start first on control plot. The start of spring phenophases on the clearcut was always observed the latest, even in comparison with the parent stand. Correlation analysis confirmed a significant correlation (P <0.05, r = -0.61) between the mean air temperature in March and April and start of the phenophase leafing in the individual years. Analysis of long-term research showed that the trend of leafing’s onset observed in course of 18 years was significant (P <0.05), manifested a shift towards earlier dates.
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