Vertebrate footprints occur in the Middle Buntsandstein (Lower Triassic) Labyrinthodontidae Beds exposed at Wióry in the northeastern margin of the Holy Cross Mountains (Poland). They represent the richest footprint assemblage from the Middle Buntsandstein in Europe known to date. This assemblage comprises 11 ichnospecies representing seven ichnogenera attributable to amphibians and reptiles. The following new ichnotaxa are erected: Prorotodactylidae ichnofam. n., Prorotodactylus mirus ichnogen. et ichnosp. n., Capitosauroides fuglewiczi ichnosp. n., Brachychirotherium wiorense ichnosp. n., Isochirotherium gierlinskii ichnosp. n., Synaptichnium kotanskii ichnosp. n., and Rhynchosauroides rdzaneki ichnosp, n. The Prorotodactylus trackmakers possibly represent a systematic group close to that from which the Rotodactylus trackmakers and dinosaurs originated.
Osteological fossils of dinosaurs are relatively rare in the Late Triassic and Early Jurassic. Thus, ichnofossils are a critical source of information on Late Triassic terrestrial vertebrate communities. The outcrops of the Tomanová Formation (?late Norian–Rhaetian) in the Tatra Mountains of Poland and Slovakia have yielded a diverse ichnofauna. Seven more or less distinct morphotypes of dinosaur tracks have been recognized and are discussed. Most tracks are partly eroded or deformed, but are preserved well enough to be assigned to a range of trackmakers, including early ornithischians, small and large theropods (coelophysoids and/or possibly early tetanurans), and probably basal sauropodomorphs (“prosauropods”) or first true sauropods.
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