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The new trench Z−17 situated in the Dębnik anticline (Cracow Region, southern Poland) exposed strata representing the Frasnian–Famennian boundary interval. The latest Frasnian crisis interval is characterized by the fauna of Ryocarhynchus tumidus interval consisting of the nominal species, Barroisella campbelli, Biernatella polonica, and representives of Lingulipora, Athyris, ?Retichonetes, Longispina, Cyrtospirifer and Warrenella. The first brachiopods that appears immediately above the F–F boundary in the survival interval include the unidentified rhipidomellid, Praewaagenoconcha cf. speciosa, and Pampoecilorhynchus geniculatus sp. nov. The fauna in the succeeding layer is dominated by P. geniculatus and Cyrtospirifer minor. The earliest Famennian repopulation assemblage consists of representatives of Barroisella, ?Rhyssochonetes, Praewaagenoconcha, Nigerinoplica, Schizophoria, unidentified rhipidomellid, Pampoecilorhynchus geniculatus sp. nov., Chapinella striata sp. nov., Crinisarina angelicoides, and Cyrtospirifer minor. Low diversity and low frequency brachiopod assemblages consisting of stunted specimens characterize the succeeding brachiopod intervals spanning the remainder of the Palmatolepis triangularis Zone. Basinal dysoxia prevailed in the region for the duration of Pa. triangularis Zone. Resumption of aerobic bottom−water conditions is marked by the appearance of brachiopods of the Dmitria gibbosa interval (Pa. crepida Zone). Resumption of favorable environmental conditions during the recovery interval is marked by an increase in brachiopod diversity recording the beginning of a strong post−extinction rediversification of the fauna.
The brachiopod faunas from deposits recording the Early–Middle Frasnian transition of Poland are poorly known. The present report describes these faunas that were recovered from Wietrznia and Kostomłoty (Holy Cross Mountains) and Dębnik (Silesian−Cracow Upland) regions positioned on the southern Polish carbonate shelf. The brachiopod distribution was analysed relative to a significant δ¹³C excursion, referred recently to as Palmatolepis punctata Event. The sporadic occurrence of brachiopods at Dębnik makes it difficult to analyse the biotic response to this geochemical anomaly. However, higher brachiopod frequency in the topmost part of the section coincides with a gradual decrease of δ¹³C down to the Frasnian background values. At two studied sections at Wietrznia the greatest taxonomic diversity and abundance of brachiopods (Flabellulirostrum–Coeloterorhynchus assemblage) are confined to the Palmatolepis transitans Zone. In the following Pa. punctata Zone brachiopods are rare (Biernatella lentiformis assemblage) or absent, which suggests a serious deterioration of the environmental conditions linked probably with periods of benthic anoxia−dysoxia and oligotrophic conditions in the basin. In two Kostomłoty sections brachiopods faunas of this age are members of the mostly monospecific rhynchonellid−dominated Phlogoiderhynchus polonicus assemblage which inhabited deeper−water environments characteristic of intrashelf oxygen−depleted basins of ŁysogóryKostomłoty region. In some horizons large shells of P. polonicus were frequently colonised by discinoid Romerella? sp. Among 28 species described in this report new forms include: Flabellulirostrum kielcensis sp. nov., F. rackii sp. nov., and Thomasaria ventosa sp. nov.
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