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Pozycja filogenetyczna owiraptorozaurow /Dinozsauria: Theropoda/

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Osmolska H., Wolsan M., Maryanska T.

Działalność Naukowa PAN. Wybrane Zagadnienia

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2002

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tom 13

51-52

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Oviraptorosaur tail forms and functions

72%

Persons,IV W. S., Currie P. J., Norell M. A.

Acta Palaeontologica Polonica

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2014

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tom 59

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nr 3

Oviraptorosaur caudal osteology is unique among theropods and is characterized by posteriorly persistent and exceptionally wide transverse processes, anteroposteriorly short centra, and a high degree of flexibility across the pre-pygostyle vertebral series. Three-dimensional digital muscle reconstructions reveal that, while oviraptorosaur tails were reduced in length relative to the tails of other theropods, they were muscularly robust. Despite overall caudal length reduction, the relative size of the M. caudofemoralis in most oviraptorosaurs was comparable with those of other non-avian theropods. The discovery of a second Nomingia specimen with a pygostyle confirms that the fused terminal vertebrae of the type specimen were not an abnormality. New evidence shows that pygostyles were also present in the oviraptorosaurs Citipati and Conchoraptor. Based on the observed osteological morphology and inferred muscle morphology, along with the recognition that many members of the group probably sported broad tail-feather fans, it is postulated that oviraptorosaur tails were uniquely adapted to serve as dynamic intraspecific display structures. Similarities, including a reduced vertebral series and a terminal pygostyle, between the tails of oviraptorosaurs and the tails of theropods widely accepted as basal members of the Avialae, appear to be convergences.

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Why tyrannosaurid forelimbs were so short: An integrative hypothesis

72%

Padian K.

Acta Palaeontologica Polonica

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2022

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tom 67

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nr 1

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An articulated pes from a small parvicursorine alvarezsauroid dinosaur from Inner Mongolia, China

72%

Hone D. W. E., Choiniere J. N., Tan Q., Xu X.

Acta Palaeontologica Polonica

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2013

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tom 58

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nr 3

A near complete and articulated parvicursorine pes from the Campanian Wulansuhai Formation is described. This pes is referred to the genus Linhenykus and is one of the first foot skeletons to be described for a derived alvarezsaur, providing new information on the first digit of the pes. The evolution of a laterally directed flange of the anterior face of the distal third metatarsal in arctometatarsalian taxa is described and discussed. This flange may have increased stability of the foot during cursorial locomotion and may also provide useful taxonomic and systematic data.

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Macroevolutionary and morphofunctional patterns in theropod skulls: A morphometric approach

72%

Foth C., Rauhut O. W. M.

Acta Palaeontologica Polonica

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2013

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tom 58

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nr 1

Theropod dinosaurs are one of the most remarkable lineages of terrestrial vertebrates in the Mesozoic, showing high taxo− nomic and ecological diversity. We investigate the cranial diversity of non−avian theropods and some basal birds, using geometric morphometrics to obtain insights into the evolutionary modifications of the skull. Theropod skulls mostly vary in the shape of the snout and length of the postorbital region (principal component [PC] 1), with further variation in orbit shape, depth of the postorbital region, and position of the jaw joint (PC 2 and PC 3). These results indicate that the cranial shape of theropods is closely correlated with phylogeny and dietary preference. Skull shapes of non−carnivorous taxa dif− fer significantly from carnivorous taxa, suggesting that dietary preference affects skull shape. Furthermore, we found a significant correlation between the first three PC axes and functional proxies (average maximum stress and an indicator of skull strength). Interestingly, basal birds occupy a large area within the morphospace, indicating a high cranial, and thus also ecological, diversity. However, we could include only a small number of basal avialan species, because their skulls are fragile and there are few good skull reconstructions. Taking the known diversity of basal birds from the Jehol biota into account, the present result might even underestimate the morphological diversity of basal avialans.

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The endocranium of the theropod dinosaur Ceratosaurus studied with computer tomography

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Sanders R. K., Smith D. K.

Acta Palaeontologica Polonica

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2005

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tom 50

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nr 3

A well preserved specimen of the theropod Ceratosaurusfrom the Upper Jurassic Morrison Formation of western Colorado was recently described and given the name C. magnicornis. The systematics of the genus is outside the scope of the present study but, as a generally accepted basal tetanuran, the braincase was CT scanned to provide a description of the endocranium, inner ear, pneumatic, and venous sinus systems in a primitive member of this clade. Five major subregions of the theropod endocranium are distinguished for the purpose of simplifying cranial computed tomographic interpretation and to provide a systematic means of comparison to other endocrania. The skull morphology of Ceratosaurus influences the overall braincase morphology and the number and distribution of the major foramina. The low pontine angle and relatively unflexed braincase is considered a more primitive character. The orientation of the horizontal semicircular canal confirms a rather horizontal and unerect posture of the head and neck. As in birds, the narrower skull morphology of Ceratosaurusis associated with fewer cranial nerve foramina. Additionally, the maxillary dominated dentigerous upper jaw of Ceratosaurusis felt to share with the alligator a large rostrally directed maxillary division of the trigeminal nerve and a small ophthalmic branch. The upper bill of birds, being dominated by the premaxillary and lacking teeth, is innervated predominantly by the ophthalmic division of the trigeminal nerve. For this reason, avian−based cranial nerve reconstructions are felt to be inappropriate for basal theropods.Ceratosaurusskull pneumatization and possible evidence of olfactory conchal structures is on the other hand very avian in character. Based on computed tomography, Ceratosaurusis determined to have possessed a typical basal theropod endocranium and bipedal vestibular system similar to Allosaurus.

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A new approach to evaluate the cursorial ability of the giant theropod Giganotosaurus carolinii

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Blanco R. E., Mazzetta G. V.

Acta Palaeontologica Polonica

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2001

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tom 46

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nr 2

The cursorial capability of the South American giant theropod Giganotosaurus carolinii should have been quite limited taking into account the strength indicator of its femur (approximately 7 GPa⁻¹) as well as the risk of experiencing grave or even lethal injuries involved in the falling of this multitonne animal on a run. However, even at low speeds a fall would have caused serious injuries. Thus, in accordance to the approach developed in this study, the maximum speed of Giganotosaurus should be not that which will implicate corporal lesions with minimum probability of lethalness. Instead, its maximum speed should be that which would permit the recovery of body equilibrium as each step is taken. Taking into consideration this approach, an indicator of stability is defined for bipedal, cursorial animals. This indicator is determined by the relationship between the time available for the movement of hip joint during the retraction of a hindlimb and the time needed to move the opposite hindlimb by an angle (in function of the speed) of sufficient magnitude as to facilitate the recovery of body equilibrium. This indicator was used to estimate the maximum speed of locomotion of Giganotosaurus (about 14 m s⁻¹) at which, from a kinematic point of view, the danger of falling does not exist.

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Avialan status for Oviraptorosauria

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Maryanska T., Osmolska H., Wolsan M.

Acta Palaeontologica Polonica

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2002

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tom 47

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nr 1

Oviraptorosauria is a clade of Cretaceous theropod dinosaurs of uncertain affinities within Maniraptoriformes. All previous phylogenetic analyses placed oviraptorosaurs outside a close relationship to birds (Avialae), recognizing Dromaeosauridae or Troodontidae, or a clade containing these two taxa (Deinonychosauria), as sister taxon to birds. Here we present the results of a phylogenetic analysis using 195 characters scored for four outgroup and 13 maniraptoriform (ingroup) terminal taxa, including new data on oviraptorids. This analysis places Oviraptorosauria within Avialae, in a sister−group relationship with Confuciusornis. Archaeopteryx, Therizinosauria, Dromaeosauridae, and Ornithomimosauria are successively more distant outgroups to the Confuciusornis−oviraptorosaur clade. Avimimus and Caudipteryx are successively more closely related to Oviraptoroidea, which contains the sister taxa Caenagnathidae and Oviraptoridae. Within Oviraptoridae, “Oviraptor” mongoliensis and Oviraptor philoceratops are successively more closely related to the Conchoraptor−Ingenia clade. Oviraptorosaurs are hypothesized to be secondarily flightless. Emended phylogenetic definitions are provided for Oviraptoridae, Caenagnathidae, Oviraptoroidea, Oviraptorosauria, Avialae, Eumaniraptora, Maniraptora, and Maniraptoriformes.

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Scapular orientation in theropods and basal birds, and the origin of flapping flight

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Senter P.

Acta Palaeontologica Polonica

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2006

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tom 51

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nr 2

Basal birds such as Archaeopteryx and Confuciusornis are typically portrayed as flapping fliers. However, here I show that shoulder joint orientation in these animals prevented elevation of the humerus above the dorsum, thereby preventing use of the recovery stroke, an important part of flapping flight. In members of the clade Ornithothoraces, which includes extant birds and the extinct avian clade Enantiornithes, the shoulder joint is reoriented to permit elevation of the humerus above the dorsum, permitting flapping flight. Although basal birds may have glided, flapping flight began significantly later in avian evolution than has been thought.

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Caudipteryx as a non-avalian theropod rather than a flightless bird

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Dyke G. J., Norell M. A.

Acta Palaeontologica Polonica

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2005

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tom 50

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nr 1

Caudipteryx zoui is a small enigmatic theropod known from the Early Cretaceous Yixian Formation of the People’s Republic of China. From the time of its initial description, this taxon has stimulated a great deal of ongoing debate regarding the phylogenetic relationship between non−avialan theropods and birds (Avialae) because it preserves structures that have been uncontroversially accepted as feathers (albeit aerodynamically unsuitable for flight). However, it has also been proposed that both the relative proportions of the hind limb bones (when compared with overall leg length), and the position of the center of mass in Caudipteryx are more similar to those seen in extant cusorial birds than they are to other non−avialan theropod dinosaurs. This conclusion has been used to imply that Caudipteryx may not have been correctly interpreted as a feathered non−avialan theropod, but instead that this taxon represents some kind of flightless bird. We review the evidence for this claim at the level of both the included fossil specimen data, and in terms of the validity of the results presented. There is no reason—phylogenetic, morphometric or otherwise—to conclude that Caudipteryx is anything other than a small non−avialan theropod dinosaur.

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Osteology of the Late Cretaceous alvarezsauroid Linhenykus monodactylus from China and comments on alvarezsauroid biogeography

58%

Xu X., Upchurch P., Ma Q., Pittman M., Choiniere J., Sullivan C., Hone D. W. E., Tan Q., Tan L., Xiao D., Han F.

Acta Palaeontologica Polonica

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2013

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tom 58

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nr 1

The alvarezsauroid theropod Linhenykus monodactylus from the Upper Cretaceous of Inner Mongolia, China is the first knownmonodactyl non−avian dinosaur, providing important information on the complex patterns of manual evolution seen in alvarezsauroids. Herewe provide a detailed description of the osteology of this taxon. Linhenykus shows a number of fea− tures that are transitional between parvicursorine and non−parvicursorine alvarezsauroids, but detailed comparisons also re− veal that some characters had a more complex distribution. We also use event−based tree−fitting to perform a quantitative analysis of alvarezsauroid biogeography incorporating several recently discovered taxa. The results suggest that there is no statistical support for previous biogeographic hypotheses that favour pure vicariance or pure dispersal scenarios as explana− tions for the distributions of alvarezsauroids across SouthAmerica, NorthAmerica andAsia. Instead, statistically significant biogeographic reconstructions suggest a dominant role for sympatric (or “within area”) events, combined with a mix of vicariance, dispersal and regional extinction. At present the alvarezsauroid data set is too small to completely resolve the biogeographic history of this group: future studies will need to create larger data sets that encompass additional clades.

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Velociraptorine dromaeosaurid teeth from the Kimmeridgian [Late Jurassic] of Germany

58%

van der Lubbe T., Richter U., Knotschke N.

Acta Palaeontologica Polonica

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2009

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tom 54

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nr 3

401-408

Six theropod teeth from a Late Jurassic (Kimmeridgian) bone bed in Langenberg Quarry of Oker (Goslar, Germany) are identified as a new dromaeosaurid taxon, here left in open nomenclature. Direct comparison reveals that the teeth are very similar to velociraptorine dromaeosaurid teeth from the Guimarota coal mine (Late Jurassic, Portugal) and to velociraptorine dromaeosaurid teeth from Uña (Barremian, Cuenca Province, Spain). Our data indicate that the teeth from the Kimmeridgian of Lower Saxony are of velociraptorine dromaeosaurid type, and therefore represent one of the oldest occurrences of the group Dromaeosauridae.

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Allometric growth in the frontals of the Mongolian theropod dinosaur Tarbosaurus bataar

58%

Yun C.-G., Peters G. F., Currie P. J.

Acta Palaeontologica Polonica

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2022

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tom 67

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nr 3

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Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics

58%

Snively E., Henderson D. M., Phillips D. S.

Acta Palaeontologica Polonica

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2006

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tom 51

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nr 3

Tyrannosaurid theropods display several unusual adaptations of the skulls and teeth. Their nasals are fused and vaulted, suggesting that these elements braced the cranium against high feeding forces. Exceptionally high strengths of maxillary teeth in Tyrannosaurus rex indicate that it could exert relatively greater feeding forces than other tyrannosaurids. Areas and second moments of area of the nasals, calculated from CT cross−sections, show higher nasal strengths for large tyrannosaurids than for Allosaurus fragilis. Cross−sectional geometry of theropod crania reveals high second moments of area in tyrannosaurids, with resulting high strengths in bending and torsion, when compared with the crania of similarly sized theropods. In tyrannosaurids trends of strength increase are positively allomeric and have similar allometric exponents, indicating correlated progression towards unusually high strengths of the feeding apparatus. Fused, arched nasals and broad crania of tyrannosaurids are consistent with deep bites that impacted bone and powerful lateral movements of the head for dismembering prey.

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Filling the ceratosaur gap: A new ceratosaurian theropod from the Early Cretaceous of Spain

58%

Sanchez-Hernandez B., Benton M. J.

Acta Palaeontologica Polonica

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2014

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tom 59

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nr 3

Ceratosaurian theropods evolved in two bursts, first in the Middle and Late Jurassic and then in the Late Cretaceous, leaving a 20 Myr gap in the Early Cretaceous during which remains are rare. We describe here a new ceratosaurian theropod, Camarillasaurus cirugedae, from fluvial deposits of the Camarillas Formation (lower Barremian, Lower Cretaceous) of Camarillas, Teruel Province, NE Spain. The new theropod is represented by a collection of associated bones, including a tooth, a possible cervical vertebra, two sternal plates, the proximal part of a right tibia, a broken right scapulocoracoid, the incomplete sacrum, five caudal vertebrae, an isolated caudal neural arch, a chevron, an almost complete presacral rib and some fragments of vertebrae, ribs, and other elements. Camarillasaurus is differentiated from other theropods by the extreme depth of the tibia proximal end, and a deep longitudinal groove on the tibia. The new dinosaur is a ceratosaur, phylogenetically close to the base of the clade, and perhaps more derived than the Chinese basal ceratosaur Limusaurus. The new taxon is significant in the evolution of the ceratosaurian dinosaurs, being placed temporally between its more common Jurassic and mid-Upper Cretaceous relatives, and it is one of only a few from Laurasia.

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An unusual theropod dinosaur from the Late Cretaceous Nemegt Formation of Mongolia

58%

Osmolska H.

Acta Palaeontologica Polonica

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1996

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tom 41

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nr 1

1-38

An incomplete skeleton of a theropod dinosaur, Bagaraatan ostromi gen, et sp. n., was found in the Nemegt Fm. at Nemegt, Mongolia. The mandible in B. ostromi has a shallow but massive dentary, relatively deep postdentary portion with two surangular foramina and somewhat elongated retroarticular process; on the lateral surface of the postacetabular process of the ilium there are two large depressions for muscle origins separated by a crestlike projection; the fibula is fused distally with the tibiotarsus and the coalesced astragalocalcaneum. Bagaraatan represents the Tetanurae and displays some synapomorphies with the Avetheropoda, however, incompleteness of the skeleton of B. ostromi does not allow to determine its more precise affiliation. Bagaraatan was about 3.0-3.5 m long, had a relatively small head and slender hind limbs. The presence of strongly developed hyposphenes in a long series of anterior caudals rendered its tail only slighily flexible proximally.

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The systematics of Late Jurassic tyrannosauroid theropods from Europe and North America

58%

Brusatte S. L., Benson R. B. J.

Acta Palaeontologica Polonica

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2013

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tom 58

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nr 1

Recent discoveries of more than ten new species of tyrannosauroid theropods are helping to understand the origin and evolu− tion of colossal body size and other characteristic features of Tyrannosaurus rex and its terminal Cretaceous relatives. Partic− ularly important has been the discovery and reinterpretation of Late Jurassic tyrannosauroids from Europe and North Amer− ica, which are intermediate in size and phylogenetic position between small basal tyrannosauroids and the largest Late Cre− taceous species. The fragmentary nature of these Jurassic specimens, however, has frustrated attempts to understand their systematics and phylogeny. A new specimen from the Late Jurassic of England was recently named as a new species (Stokesosaurus langhami) of the genus Stokesosaurus, which is known from several fragmentary fossils from North Amer− ica. We review the systematics and phylogeny of these European and North American specimens and show that there are no unequivocal synapomorphies uniting them. Furthermore, a revised phylogenetic analysis does not recover them as sister taxa. This necessitates a taxonomic revision of this material, and we name a new genus (Juratyrant) for theBritish specimen.

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New information on scavenging and selective feeding behaviour of tyrannosaurids

58%

Hone D. W. E., Watabe M.

Acta Palaeontologica Polonica

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2010

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tom 55

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nr 4

Feeding traces for carnivorous theropod dinosaurs are typically rare but can provide important evidence of prey choice and mode of feeding. Here we report a humerus of the hadrosaurine Saurolophus which was heavily damaged from feeding attributed to the giant tyrannosaurine Tarbosaurus. The bone shows multiple bites made in three distinctive styles termed “punctures”, “drag marks” and “bite−and−drag marks”. The distribution of these bites suggest that the animal was actively selecting which biting style to use based on which part of the bone was being engaged. The lack of damage to the rest of the otherwise complete and articulated hadrosaur strongly implies that this was a scavenging event, the first reported for a tyrannosaurid, and not feeding at a kill site.

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Variation in premaxillary tooth count and a developmental abnormality in a tyrannosaurid dinosaur

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Miyashita T., Tanke D. H., Currie P. J.

Acta Palaeontologica Polonica

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2010

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tom 55

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nr 4

Premaxillary tooth count tends to be stable amongst toothed dinosaurs, and most theropods have four teeth in each premaxilla. Only one case of bilaterally asymmetric variation is known in theropod premaxillary dentition, and there is no record of ontogenetic or individual variation in premaxillary tooth count. Based on these observations, a tyrannosaurid left premaxilla with three teeth (TMP 2007.20.124) is an interesting deviation and represents an unusual individual of Daspletosaurus sp. with a developmental abnormality. The lower number of teeth is coupled with relatively larger alveoli, each of which is capable of hosting a larger than normal tooth. This indicates that tooth size and dental count vary inversely, and instances of reduction in tooth count may arise from selection for increased tooth size. On the other hand, the conservative number of premaxillary teeth in most theropods implies strong developmental constraints and a functional trade−off between the dimensions of the premaxillary alveolar margin and the size of the teeth. In light of recent advances in the study of tooth morphogenesis, tooth count is a function of two parameters: dimensions of an odontogenic field for a tooth series, and dimensions of tooth positions. A probable developmental cause for the low tooth count of TMP 2007.20.124 is that the dimensions of the alveoli expanded by approximately a third during tooth morphogenesis. Numerical traits such as tooth count are difficult to treat in a phylogenetic analysis. When formulating a phylogenetic character, a potential alternative to simply counting is to rely on the morphological signature for developmental parameters that control the number of the element in question.

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Isolated theropod teeth from the Middle Jurassic of Niger and the early dental evolution of Spinosauridae

58%

Serrano-Martínez A., Vidal D., Sciscio L., Ortega F., Knoll F.

Acta Palaeontologica Polonica

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2016

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tom 61

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nr 2

Four isolated theropod teeth from the ?Bathonian “Argiles de l’Irhazer” in Niger are described. The teeth were found in association with the holotype of the basal sauropod Spinophorosaurus nigerensis. These specimens have been assigned to two different taxa by independent analyses, such as direct comparison with teeth previously described in the literature, discriminant and morphometric analyses from metric characters, and cladistic and cluster analyses from discrete characters. The results suggest that three teeth share affinities with those of Megalosauridae and Allosauridae, belonging most likely to the former. The fourth tooth might be from a member of the stem group Spinosauridae. If so, this would be the oldest representative of this clade. This tooth shows a combination of characters that are unusual in typical spinosaurid teeth (crown moderately compressed labiolingually and curved distally with minute denticles on the carina and a deeply veined enamel surface texture without apicobasal ridges). This could shed light on the morphological transition from the plesiomorphic ziphodont dental pattern to that of Spinosauridae. This tooth would also allow a better understanding of the origin of the spinosaurids, supporting a Gondwanan origin for the group.

Ograniczanie wyników

Pozycja filogenetyczna owiraptorozaurow /Dinozsauria: Theropoda/

Oviraptorosaur tail forms and functions

Why tyrannosaurid forelimbs were so short: An integrative hypothesis

An articulated pes from a small parvicursorine alvarezsauroid dinosaur from Inner Mongolia, China

Macroevolutionary and morphofunctional patterns in theropod skulls: A morphometric approach

The endocranium of the theropod dinosaur Ceratosaurus studied with computer tomography

A new approach to evaluate the cursorial ability of the giant theropod Giganotosaurus carolinii

Avialan status for Oviraptorosauria

Scapular orientation in theropods and basal birds, and the origin of flapping flight

Caudipteryx as a non-avalian theropod rather than a flightless bird

Osteology of the Late Cretaceous alvarezsauroid Linhenykus monodactylus from China and comments on alvarezsauroid biogeography

Velociraptorine dromaeosaurid teeth from the Kimmeridgian [Late Jurassic] of Germany

Allometric growth in the frontals of the Mongolian theropod dinosaur Tarbosaurus bataar

Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics

Filling the ceratosaur gap: A new ceratosaurian theropod from the Early Cretaceous of Spain

An unusual theropod dinosaur from the Late Cretaceous Nemegt Formation of Mongolia

The systematics of Late Jurassic tyrannosauroid theropods from Europe and North America

New information on scavenging and selective feeding behaviour of tyrannosaurids

Variation in premaxillary tooth count and a developmental abnormality in a tyrannosaurid dinosaur

Isolated theropod teeth from the Middle Jurassic of Niger and the early dental evolution of Spinosauridae