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Jurassic rhynchonellide brachiopods from the Jordan Valley are herein revised and new taxa are added to the faunal list. In this study of Jurassic rhynchonellides from Wadi Zarqa, northwestern Jordan, we recognize the following taxa: Eurysites rotundus, Cymatorhynchia quadriplicata, Daghanirhynchia triangulata, D. angulocostata, Pycnoria magna, Schizoria elongata, and Schizoria cf. intermedia. The following new taxa are described: Daghanirhynchia susanae sp. nov. and Amydroptychus markowitzi sp. nov. The Middle Jurassic Mughanniyya Formation of northwest Jordan is dominated by limestone beds. The sedimentary environment is interpreted as neritic, light, and nutrient−rich resulting in high faunal diversity. The high rhynchonellide endemism of this fauna is yet another confirmation of pronounced Middle Jurassic endemism along the southern Tethyan margin of the Ethiopian Province. Brachiopods of the Jordanian Mughanniyya Formation can be correlated with the fauna of the Aroussiah Formation in Sinai and the Zohar and Matmor formations in Southern Israel.
The rhynchonellid species, Pammegetherhynchus kowalaensis sp. n., occurs in the late Frasnian (Early to Late Palmatolepis rhenana, and possibly early Palmatolepis linguiformis conodont zones) marly-bituminous succession at Kowala (various outcrops) in the Gałęzice Syncline, south of Kielce in the Holy Cross Mountains, Poland. The only other known species of this genus is the type species, Pammegetherhynchus merodae Sartenaer, 1977, from the late Frasnian (somewhere in the Early and Late Palmatolepis rhenana Zones) of the French Fagne (dark shales of 'Matagne' aspect), and, probably, of the Eifel ('Büdesheimer Goniatitenschiefer'). P. kowalaensis sp. n. occurred in level-bottom pioneer assemblages, thriving in reef downslope, mostly poorly-oxygenated habitats of the Kellwasser interval. The species finally disappeared near the Frasnian-Famennian boundary. The genus Pammegetherhynchus seems to be particularly suited to stressed deep-water shelf environments in the European part of the Laurussian shelf, widely distributed in this crisis time.
In contrast to dramatic losses of the Atrypida and Pentamerida at the Frasnian–Famennian boundary, the Rhynchonellida demonstrated relatively high rate of survival, and recolonized vacated benthic ecospace after the F–F extinction events. The Late Devonian evolution of rhynchonellid faunas from the East European Platform shows three distinctive periods of their mass appearance. High abundance of rhynchonellids is characteristic for early–middle Frasnian (Palmatolepis transitans–Pa. punctata zones) and early Famennian (Pa. crepida Zone). Invasion of taxonomically diverse and rich rhynchonellid faunas usually corresponds to the major transgressive episodes, whereas decline coincides with regressive conditions of the basin. Rhynchonellid assemblages were replaced in the late Frasnian (Late Pa. hassi–Pa. linguiformis zones) by theodossiid− and cyrtospiriferid−dominated assemblages, which occupied habitats in newly expanding marine environments. The extinction of theodossiids at the end of the Frasnian and the next transgressive episode possibly stimulated an expansion of rhynchonellids. The early Frasnian species Ripidiorhynchus livonicus (Buch, 1834), and the early Famennian R. huotinus (Verneuil, 1845) and R. griasicus (Nalivkin, 1934) are revised. Early Famennian species Paromoeopygma koscharica (Nalivkin, 1934) from the central region is redescribed. Ripidiorhynchus chencinensis sp. nov. from the latest Givetian of Poland, as well as Globulirhynchia minima sp. nov. from the late Frasnian of the central region of the Russia, are described.
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