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Fallomyrma, a new monotypie ant genus from the Rovno, Saxonian, and Danish ambers (Late Eocene), and a new species, F. transversa, are described. The taxonomic position and morphological similarity of the new genus to other genera is discussed.
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A new ant genus from the late Eocene European amber

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Eocenomyrma gen. nov. of extinct ants of the family Formicidae, subfamily Myrmicinae, is described from the late Eocene European amber (ca. 40 Ma), based on six specimens from six pieces of amber; three of them contain E. rugosostriata (Baltic and Saxonian ambers); the remainder contain three new species: E. orthospina (Baltic Amber), E. electrina (Scandinavian Amber), and E. elegantula (Baltic Amber). Eocenomyrma resembles two extant genera: Myrmica and Temnothorax (both of which also occur in late Eocene European amber), but differs from them by the following apomorphies: clypeus short and broad, with two lateral longitudinal carinae and distinctly marked anterolateral corners, its median portion faintly concave transversally, anterior margin broad and shallowly concave medially, with pairs of long setae situated on the anterolateral clypeal corners, and central part of the anterior clypeal margin without setae; middle and hind tibiae lacking the spurs. Palp formula in Eocenomyrma is 4, 3 versus 6, 4 in Myrmica. We include Eocenomyrma in the tribe Formicoxenini. Nothomyrmica rugosostriata is transferred to Eocenomyrma, and the neotype of the latter species is designated; Nothomyrmica petiolata is transferred to the genus Temnothorax. A key for the identification of all known Eocenomyrma species is compiled.
The extinct ant genera Stigmomyrmex Mayr and Stiphromyrmex Wheeler (subfamily Myrmicinae) were described from the Baltic amber (Late Eocene, Priabonian, ca. 34-38 Ma). A new species, Stigmomyrmex rugulosus sp. nov., is described from the Baltic and Bitterfeld ambers, and a neotype of Stiphromyrmex robustus (Mayr) is designated. Additionally, we found from the Bitterfeld amber three more specimens of S. venustus Mayr, previously the only known species of the genus. S. rugulosus well differs from S. venustus by the presence of longitudinal rugulosity on the head dorsum, mesosoma and waist, and by the presence of numerous suberect hairs on the legs.
Tetramorium galaticum Menozzi, 1936, new status and Tetramorium hippocratis Agosti & Collingwood, 1987, members of Tetramorium semilaeve complex, are redescribed based on new materials from Greece and western Turkey. Detailed description of gyne and male is given for the first time. Two new species of this complex with all castes are described: Tetramorium kephalosi (Greece and Croatia) and Tetramorium bellerophoni (Turkey). Differential diagnoses from T. semilaeve, colour photos of all castes and male genitalia are given.
Two extinct genera of ants from the late Eocene (ca. 40 Ma), Protomyrmica gen. nov. and Plesiomyrmex gen. nov. (family Formicidae, subfamily Myrmicinae), are described based on single specimens (males), from Baltic and Bitterfeld (also called Saxonian) ambers respectively; both genera belong to the tribe Myrmicini. In gross morphology they are similar to modern Myrmica but have a series of apomorphies combined with characters that are plesiomorphic not only in the tribe Myrmicini, but also in the subfamily Myrmicinae. The most significant plesiomorphies concern the antennal structure and wing venation of both genera. The antennal scape is short and the funiculus is filiform, having no apical club. Moreover, the antennae of Protomyrmica are “sphecoid” with the length of the funicular segments gradually decreasing towards the apex (i.e., the longest is basal, starting from the second, and the shortest is apical); this type of structure is basal for the family Formicidae as a whole. Although we consider the wing venation of Protomyrmica to represent the prototype of wings in the subfamily Myrmicinae, it has an apomorphy absent in the modern Myrmicini genera—the antennae are inserted into the head well behind the posterior margin of the clypeus. Plesiomyrmex also has a peculiar apomorphy not found in any other genus of Myrmicinae: the antennae are inserted into toruli located on short sub−vertical tube−like or cup−like structures that protrude distinctly above the head surface. As a result, we do not consider either of the newly described genera to be the direct ancestors of modern Myrmicini; nevertheless, the presence of very ancient plesiomorphies may indicate their antiquity, and thus the latest estimated time for the origin of the tribe Myrmicini should be at least the early Eocene.
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