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Sont présentés les résultats des observations sur les formations récifales dans le Frasnien et le Givétien des Monts de Sainte-Croix (Góry Świętokrzyskie), dans les localités Kadzielnia, Sitkówka et Jurkowice. A Kadzielnia ce sont les biohermes et à Sitkówka et Jurkowice - les biostromes.
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W obrębie podrzędu Phillipsastraeacea autorka ustanawia nową rodzinę Marisastridae, obejmującą formy kolonijne o masywnych koloniach, bądź formy osobnicze. Cechą charakterystyczną nowej rodziny jest obecność wyraźnej epiteki, wrzecionowatych septów dwojakiej długości, złożonych tabul oraz brak podkówek. Do nowej rodziny zaliczono Marisastrum n.gen. i Ceratophyllum Gürich, 1896.
New chondrichthyan microremains from several Frasnian–Famennian sections in the Holy Cross Mountains and Dębnik area (Southern Poland) are investigated and compared to previous data. The reaction of different groups of chondrichthyans to environmental changes during the Kellwasser Event is analysed. Following the extinction of phoebodont sharks of Phoebodus bifurcatus group before the end of the Frasnian, only two chondrichthyan species, viz. Protacrodus vetustus Jaekel, 1921 and Stethacanthus resistens sp. nov. (possibly closely related to “Cladodus” wildungensis Jaekel, 1921), occur in the upper part of Frasnian Palmatolepis linguiformis conodont Zone and persist into the Famennian. Global cooling is considered a possible cause of the extinction of Frasnian subtropical phoebodonts on Laurussian margins.
Retained colour pattern on the shells of Plectodonta sp. from the earliest Devonian of Podolia (Ukraine) is the first finding for strophomenide brachiopods and the oldest among articulate brachiopods. The colour pattern in Plectodonta sp. is composed of small, round, brownish spots scattered rather irregularly on the ventral valve only. This may suggest that the described pattern probably performed a protective function through disruptive camouflage against visual systems of potential predators. The occurrence of the colour pattern in Plectodonta sp. exclusively on the ventral valve strongly suggests that these brachiopods lived with the patterned (and convex) ventral valve upwards and the patternless concave dorsal valve facing to the underlying substrate. It thus contradicts a general assumption that concavo−convex brachiopods lived with their convex valves resting on the sediment.
In the Podolian Dniester Basin (southwestern Ukraine) the Lower Devonian marine deposits are represented by about 530 m thick continuous sequence of interlaminated carbonate and schale outcrops at several localities. Conodonts occur in most of the carbonate layers of the whole Lochkovian but are not abundant and their ramiform elements are mostly broken or lacking. Therefore, only the pectiniform, Pa elements of twenty five stratigraphically important conodont species occurring in the region are discussed and two new species, Caudicriodus schoenlaubi and Pandorinellina? parva are proposed. The hypothetical phyletic relationships within the main representatives of the icriodontid and spathognathodontid genera, Caudicriodus, Zieglerodina, and Pandorinellina? are traced. Comparison of the previously published and newly obtained data revealed discrepancies in the hitherto used interpretation of some of the conodont taxa and their stratigraphic ranges. Contrary to the earlier reports, Caudicriodus postwoschmidti does not occur in the lower Lochkovian but only in the middle part of the Chortkiv Formation, high above the Monograptus uniformis Zone. Based on new material and verification of the previous determinations, a modified scheme of the Lochkovian conodont zonation in Podolia is proposed. Conodont zones: Caudicriodus hesperius, C. transiens, C. postwoschmidti, C. serus, and ?Caudicriodus steinachensis are distinguished. The zones are correlated with conodont zonations in other regions—Barrandian, Cantabrian Mountains, Pyrenees, and Nevada. Biostratigraphy of the Siluro−Devonian transition and Lochkovian is integrated with the carbon isotope stratigraphy.
Ischnacanthiform acanthodian dentigerous jaw bones from the Lower Devonian (Late Lochkovian) of Podolia are described for the first time. One new genus and one new species are established. Podoliacanthus gen. nov. is diagnosed as having small−sized jaw bones, the presence of specific accessory cusps/denticles on the medial side of teeth of the lateral tooth row, and groups of denticles forming the lingual tooth row. Podoliacanthus zychisp. nov. is distinguished in having elongated slender jaw bones and lateral teeth with one medial side denticle. Besides, three species are described in open nomenclature: Podoliacanthus sp. 1, while similar to Podoliacanthus zychi sp. nov., differs in having stronger posterior inclination of the teeth tips and presence of well developed flanges of the teeth, P. sp. 2 has quite robust jaw bones and teeth with two medial side denticles, and Podoliacanthus sp. 3 has small narrow jaw bones and teeth with three medial side denticles. Morphology of the lingual tooth row is considered to be a diagnostic feature of generic and higher taxonomic levels, while accessory medial cusps/denticles of the teeth are regarded as diagnostic features at species level. The new genus also occurs in Upper Silurian or Lower Devonian deposits of North Greenland. Preservation of the jaw bones possibly depends on their secondary mineralization.
Investigation of mixed carbonate−siliciclastic Lower Devonian deposits have been carried out in the Ivanye Zolote and Ustechko sections in Podolia, Ukraine. Based on palynomorph evidence, the age of the samples studied is late Lochkovian, not older than the NM Oppel Miospore Zone, specifically the Si Lineage Zone. The presence of acritarchs and chitinozoans points to dominantly marine depositional conditions. However, a regressive environmental change toward more brackish conditions is indicated by a decrease in the taxonomic diversity of acritarchs in the topmost samples, the simultaneous disappearance of chitinozoans, and an increase in leiosphaerid frequency. Furthermore, evolution of limestone microfacies demonstrates a progressive transition from a shrinking marine basin toward a brackish, storm−affected muddy lagoon, manifested by recurrent profusion of impoverished, mostly opportunistic and euryhaline shelly benthos (nuculanid bivalves, leperditicopids and other ostracods, terebratulid brachiopods), chaetetid demosponges and diverse ichthyofauna. The association of plant (mainly nematophytes and some tracheids) and animal (eurypterid, ?scorpion, and possibly other arthropod) remains points to the presence of nearby Early Devonian wetland vegetation, providing food and shelter for various semi−aquatic and other terrestrial arthropods.
Nine species and 3 subspecies of the Couvinian and Givetian bryozoans of the order Ctenostomata Busk, belonging to 5 genera: Ropalonaria Miller, Vinella Ulrich, Allonema Ulrich & Bassler, Ascodictyon Nicholson & Etheridge, Jr, and Eliasopora Bassler are described. Two of the described species have not been given names (Vinella sp. and Ascodictyon sp.), one is a new subspecies (Allonema moniliforme parvum n. subsp.) and five - new species (Ropalonaria givetiana n. sp., Ascodictyon sparsiforme n. sp., Ascodictyon vinelliforme n. sp., Ascodictyon venustum n. sp. and Eliasopora devoniana n. sp.). The specimens in vestigated come from the localities Skały and Grzegorzowice, situated in the eastern part of the Bodzentyn syncline. The problems of age, stratigraphic and geographical range, as well as ecological conditions of the Ctenostomata assemblage investigated are discussed.
Six species, of this number 2 new ones, and a new subspecies, all of them belonging to the family Lindstroemiidae Počta, as well as one species of the family Amplexocariniidae Różk., from the Lower-Givetian brachiopod shales from Skały, Holy Cross Mountains (Góry Świętokrzyskie), Poland, are described.
In the classic section across the Silurian–Devonian boundary at Dnistrove (Podolia, Ukraine) the brachiopod fauna has never been studied in detail. This paper presents results of research on brachiopods from this important locality and time interval. Bed−by−bed collecting has enabled the detailed distribution of brachiopod taxa through the boundary beds to be revealed. Generally, the reference section at Dnistrove yields rather scarce but often well preserved brachiopods. Dayia bohemica and Dnestrina gutta can be regarded as characteristic species for the uppermost Silurian. A relatively high−diversity but low−abundance brachiopod fauna occurs in the lowest 1.8 m of the earliest Devonian. Only three forms have been found to cross the Silurian–Devonian boundary: the strophomenide Plectodonta (Plectodonta) mariae pantherae subsp. nov., the atrypide Gracianella (Sublepida) paulula sp. nov., and the spiriferide Howellella (Howellella) latisinuata. A relatively narrow brachiopod−rich interval at 5.5 m above the Silurian–Devonian boundary yields 16 brachiopod species which probably indicate a setting near the lower limit of the photic zone equivalent to the Benthic Assemblage 3–4 boundary. Two new species and one new subspecies are described: Skenidioides tatyanae, Plectodonta (Plectodonta) mariae pantherae, and Gracianella (Sublepida) paulula.
The first appearance of the brachiopod Cyrtospirifer and related forms in the Late Devonian of South China significantly postdates the beginning of the Frasnian and the entry of the group in other parts of the world. Scattered data from different sections suggest that its first entry, associated with the emergence of other plicate spiriferids, such as theodossid and conispiriferid brachiopods, was late in the Middle Frasnian. At the same time, many rhynchonellids disappeared or became extinct locally in South China. This brachiopod faunal overturn near the Palmatolepis punctata–Early Pa. hassi zonal boundary is the most significant event in the Early–Middle Frasnian of South China, characterized by about a 35% loss of existing species and the flourishing of the plicate spiriferids, which was coeval with the end of a major biogeochemical perturbation recently recognized in the Pa. punctata Zone. By contrast, atrypid brachiopods do not seem to show any significant diversity change. The brachiopod faunal change was probably related to a (local?) transgressive event in South China, which also brought new pelagic faunas northwards into some intra−shelf deeper water areas, such as the Shetianqiao area in central Hunan Province. Fifteen brachiopod species are described and illustrated, which include some taxa that are first recorded or recognized in South China, e.g., the spiriferid Pyramidaspirifer, which is now known from both North America and South China. One new species, Desquamatia qiziqiaoensis, is erected.
The fossil Charophyta from the Devonian sediments of Spitsbergen are here described for the first time and classified to the species rank on the basis of thin sections.
During the Devonian the sedimentation on the continental shelves of Ardenne Massif and Boulonnais has changed from a mixed siliciclastic-carbonate ramp (Eifelian), through a carbonate barrier reef (Givetian) and then to a detritic influx with local mud-mounds (Frasnian). Here we analysed the faunistic dynamics of the trilobite associations through the changing environment. We used multivariate analyses (clustering and ordering) to discriminate the trilobite associations within 67 different samples. Three previously known communities and one new were recognised: the Eifelian Mixed association, the Givetian Dechenella association and the two Frasnian Bradocryphaeus and Scutellum-Goldius associations. These trilobite faunas present a progressive ecological specialisation. The Mixed association occurs both in the ramp or carbonated (local reef developed on the ramp) facies without any significant difference in its composition. The Dechenella fauna occurs preferentially close to barrier reefs, but can also survive during short periods of detrital input. The two Frasnian communities show a strong relationship with their environment. The Scutellum-Goldius association is only found in reef systems, whereas the Bradocryphaeus flourishes exclusively in lateral facies.
Whereas cosmopolitan distribution patterns are established for many Late Devonian vertebrates (e.g., placoderms, onychodontiforms), few palaeobiogeographic studies have considered chondrichthyans. Recent discoveries of shark material demonstrate that some chondrichthyans were cosmopolitan by the Middle Devonian. Abundant Givetian microremains have been recovered from the Cairo quarry in eastern New York State, USA. These include teeth of two shark species with Gondwanan affinities, the omalodontid Portalodus mannoliniae sp. nov. and the antarctilamnid Wellerodus priscus. Abundant teeth of P. mannoliniae sp. nov. are characterized by a smooth diplodont crown, polarized cusps, and a labially oriented base. The teeth demonstrate monognathic heterodonty. The juvenile morph is distinguished from the adult by smaller size, slender cusps, and variation in the shape of the base. W. priscus is represented by rare juvenile teeth. Two groups of scales that show affinity to material from northern (Spain) and East Gondwana (Antarctica) are tentatively attributed to the two described species. Antarctilamnid distribution suggests a north Gondwanan origin and a colonization of the margin of the landmass before dispersing to Laurentia by the Middle Devonian. This material further indicates that vertebrate global dispersal was initiated by the Middle Devonian, and emphasizes earlier palaeogeographic interpretations that the Middle Devonian “Hamilton fauna” of North American Laurussia originated in the Early Devonian in South American Gondwana.
Anomalous development of shell ornamentation and repaired shell injury in the Early Devonian dacryoconarid tentaculites are documented and interpreted as either a repaired injury of the shell (evidence of unsuccessful predation obscured by recrystallization), or as a result of an anomalous function of the mantle, caused by injury of the soft body. The manner of shell repair, which resembles the way that some modern marine animals, such as mollusks, repair their shells, is discussed. The issue of phylogenetic affinities of tentaculites has been also outlined. These findings represent the first documentation of unsuccessful predation on the Middle Paleozoic plankton.
The well−known association of platyceratid snails and crinoids typically involves a single snail positioned on the tegmen of the crinoid host; this has led to the inference of coprophagy. Two specimens of the camerate crinoid Arthroacantha from the Middle Devonian Silica Formation of Ohio, USA, exhibit numerous snails on their tegmens. On one of these, 6 platyceratid juveniles of approximately equal size are found on the tegmen. On the second crinoid, the largest of 7 infesting platyceratids occupies the typical position over the anal vent while others are either superposed (tiered) upon it or are positioned elsewhere on the tegmen. These specimens illustrate that platyceratids (1) settled on crinoids as spat, (2) were not strictly coprophagous during life yet (3) benefited from a position over the anal vent.
A new discinoid brachiopod Gigadiscina gen. nov., with the type species G. lessardi sp. nov., is described from the Lower Devonian (Siegenian) of the Tamesna Basin (South Ahaggar Massif, South Algeria). It is characterised by large size and convexo−planar profile of the shell, with a subcentral pedicle foramen. Micro−ornament is typically discinoid, with small circular pits in radial rows on the post−larval shell surface. Related species of Malvinokaffric Realm origin from South Africa, Falkland Islands, Antarctica, South America, and Libya are reviewed, including the poorly known Discina anomala from the Lower Devonian of Germany. The giant size and convexo−planar shells of these discinoids, remarkably similar to recent limpets, are interpreted as adaptation to a habitat in proximity of sandy and gravel beaches in a high−energy environment. Most likely, the conical dorsal valve suppressed drag in turbulent waters, whereas fixation of shell by large, sucker−like pedicle eliminated peeling from the substrate.
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A new Middle Devonian heterocoral from Spain

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Most of the heterocorals described up to now from strata older than Famennian are of doubtful affinities. Numerous specimens of the new genus and new species here described were collected during a sedimentological study of the Santa Lucía Formation in La Pola de Gordon (León province, Cantabrian Mountains, NW Spain). Stellaphyllia luciensis gen. et sp. nov. is characterized by cylindrical coralla, with a diameter of less than 2 mm and more than 2 cm long, having stellate to more or less rounded outer profile. Four to six septa, less frequently seven, are cojoined axially by a short oblique septum. The few tabulae present are steeply arched. The microstructure of the septa and interseptal elements shows thin elongate microlamellae parallel to a median plate which is granular in appearance. The new genus dated to the lower Eifelian age could be important in understanding the early phylogeny and origin of the order Heterocorallia.
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