Phylogenetic position of the crocodylian Megadontosuchus arduini and tomistomine palaeobiogeography

• Autorzy: Piras P , Delfino M. , Del Favero L. , Kotsakis T.
• Języki publikacji: EN

Abstrakty

EN

A cladistic analysis of Megadontosuchus arduini from the middle Eocene of Monte Duello (NE Italy) confirms its tomistomine relationships, but the low number of scorable characters determines a low resolution within the tomistomine clade. However, Megadontosuchus is clearly distinct from the other Eocene European or North African tomistomines, in having a moderate elongated but robust rostrum, massive maxillary and dentary teeth and large supratemporal fenestrae. The rostrum and teeth characteristics could indicate that M. arduini had a degree of feeding specialization intermediate between Maroccosuchus zennaroi and the Eocene European tomistomines. A summary of tomistomine palaeobiogeography suggests that despite only one species with a rather restricted range survives at present, such a clade had a glorious past with a world wide distribution documented by a conspicuous fossil record that starts at least in the early Eocene. At present, a detailed knowledge of tomistomine palaeobiogeography is hindered by the lack of modern taxonomic revisions of some fossil remains and therefore by the poor understanding of phylogenetic relationships.

Słowa kluczowe

EN

Tomistominae; Eocene; Italy; Tomistoma schlegelii; Megadontosuchus arduini; phylogenetic position; Crocodylidae; paleobiogeography; paleontology

• Wydawca: -
• Czasopismo: Acta Palaeontologica Polonica
• Rocznik: 2007
• Tom: 52
• Numer: 2
• Opis fizyczny: p.315-328,fig.,ref.

Twórcy

autor

Piras P

• Universita Roma Tre, Largo S.Leonardo Murialdo 1, I-00146, Roma, Italy

autor

Delfino M.

autor

Del Favero L.

autor

Kotsakis T.

Bibliografia

• Agustí, J. and Antón, B. 2002. Mammoths, Sabretooths and Hominids: 65 Million Years of Mammalian Evolution in Europe. 313 pp. Columbia Univ. Press, New York.
• Aldinio, P. 1896. Sul Tomistoma (Gavialosuchus) lyceensis del calcare miocenico di Lecce. Atti dell’Accademia Gioenia di Scienze Naturali in Catania (Serie 4) 9: 1–11.
• Altichieri, L. 1980. Il giacimento di Monte Duello (Verona). In: I vertebrati fossili italiani. Catalogo delle Mostra. Verona 1980, 159–160. Museo Civico di Storia Naturale, Verona.
• Andrews, C.W. 1901. Preliminary note on some recently discovered extinct vertebrates from Egypt. Geological Magazine, Decade 4 8: 436–444.
• Andrews, C.W. 1905. Notes on some new Crocodilia from the Eocene of Egypt. Geological Magazine, Decade 5 2: 481–484.
• Antunes, M.T. 1961. Tomistoma lusitanica, crocodilien du Miocène du Portugal. Revista da Facultade de Ciências Universidad de Lisbõa, Serie IIc 9: 3–88.
• Antunes, M.T. 1987. Affinities and taxonomical status of Miocene longirostrine crocodilians from Western Europe with remarks on phylogeny, paleoecology and distribution. Comunicaçoes dos Serviços Geológicos de Portugal 73: 49–58.
• Antunes, M.T. 1994. On Western Europe Miocene gavials (Crocodylia) their paleogeography, migrations and climatic significance. Comunicaçoes dos Serviços Geológicos de Portugal 80: 57–69.
• Antunes, M.T. and Cahuzac, B. 1999. Crocodilian faunal renewal in the Upper Oligocene of Western Europe. Comptes Rendus de l’Académie des Sciences de Paris, Série 2 328: 67–63.
• Antunes, M.T. and Ginsburg, L. 1989. Les crocodiliens des faluns miocènes de l’Anjou. Bulletin du Musée Nationale d’Histoire naturelle de Paris, Section C 11: 79–99.
• Aoki, R. 1983. A new generic allocation of Tomistoma machikanense, a fossil crocodilian from the Pleistocene of Japan. Copeia 1983: 89–95.
• Aoki, R. 1992. Fossil crocodilians from the Late Tertiary strata in the Sinda Basin, Eastern Zaire. African Study Monographs, Supplement 17: 67–85.
• Arambourg, C. and Magnier, P. 1961. Gisements de vertébrés dans le bassin tertiaire de Syrte (Libye). Comptes Rendus de l’Académie des Sciences de Paris, Série D 252: 1181–1183.
• Astre, G. 1931. Les crocodiliens fossiles des terrains tertiaires sous−Pyrénéens. Bulletin de la Société d’Histoire naturelle de Toulouse 61: 25–71.
• Auffenberg, W. 1954. Additional specimens of Gavialosuchus americana (Sellards) from a new locality in Florida. Quarterly Journal of Florida Academy of Sciences 17: 185–209.
• Benton, M.J. and Clark, J.M. 1988. Archosaur phylogeny and the relationships of the Crocodilia. In: M.J. Benton (ed.), The Phylogeny and Classification of the Tetrapods. Vol. 1, 295–338, Clarendon Press, Oxford.
• Berg, D.E. 1969. Charactosuchus kugleri, von Jamaica. Eclogae Geologicae Helvetiae 62: 731–735.
• Brochu, C.A. 1996. Closure of neurocentral sutures during crocodilian ontogeny: implications for maturity assessment in fossil archosaurs. Journal of Vertebrate Paleontology 16: 49–62.
• Brochu, C.A. 1997. Phylogenetic Systematics and Taxonomy of Crocodylia. Unpublished Ph.D. thesis. 467 pp. University of Texas, Austin.
• Brochu, C.A. 1999. Phylogeny, systematics, and historical biogeography of Alligatoroidea. Society of Vertebrate Paleontology Memoirs 6: 9–100.
• Brochu, C.A. 2001. Congruence between physiology, phylogenetics, and the fossil record on crocodylian historical biogeography. In: G. Grigg, F. Seebacher, and C.E. Franklin (eds.), Crocodilian Biology and Evolution, 9–28. Surrey Beatty and Sons, Sydney.
• Brochu, C.A. 2003. Phylogenetic approaches toward crocodylian history. Annual Review of Earth and Planetary Science 31: 357–397.
• Brochu, C.A. 2004. A new Late Cretaceous gavialoid crocodylian from eastern North America and the phylogenetic relationships of Thoracosaurs. Journal of Vertebrate Paleontology 24: 610–633.
• Brochu, C.A. 2006. Osteology and phylogenetic significance of Eosuchus minor (Marsh 1870), new combination, a longirostrine crocodylian from the Late Paleocene of North America. Journal of Paleontology 80: 162–186.
• Brochu, C.A. (in press). Systematics and taxonomy of Eocene tomistomine crocodylians from Britain and Northern Europe. Palaeontology.
• Brochu, C.A. and Gingerich, P.D. 2000. New tomistomine crocodylian from the middle Eocene (Bartonian) of Wadi Hitan, Fayum Province, Egypt. Contributions of Museum of Paleontology of University of Michigan 30: 251–268.
• Brochu, C.A. and Rincón, A.D. 2004. A gavialoid crocodylian from the Lower Miocene of Venezuela. In: M. Sánchez−Villagra and J. Clack (eds.), Fossils of the Castillo Formation, Venezuela: Contributions in Neotropical Palaeontology. Special Papers in Palaeontology 71: 61–79.
• Broin, F. de 1988. Les tortues de Gondwana. Studia Palaeocheloniologica 2: 103–142.
• Buckland, W. 1836. Geology and Mineralogy Considered with Reference to Natural Theology. 618 pp. Pickering, London.
• Buffetaut, E. 1982. Systématique, origine et évolution des Gavialidae sud−américains. Geobios, Mémoire spéciale 6: 127–140.
• Buffetaut, E., Crouzel, F., Juillard, F., and Stigliani, F. 1984. Le crocodilien longirostre Gavialosuchus dans le Miocène moyen de Polastron (Gers, France). Geobios 17: 113–117.
• Buscalioni, A.D., Ortega, F., Weishampel, D.B., and Jianu, C.M. 2001. A revision of the crocodyliform Allodaposuchus precedens from the Upper Cretaceous of the Hateg Basin, Romania. Its relevance in the phylogeny of Eusuchia. Journal of Vertebrate Paleontology 21: 74–86.
• Capellini, G. 1890a. Sul coccodrilliano garialoide (Tomistoma calaritanus) scoperto nella collina di Cagliari nel 1868. Rendiconti della Reale Accademia dei Lincei, serie 4 6 (I semestre): 149–151.
• Capellini, G. 1890b. Sul coccodrilliano garialoide (Tomistoma calaritanus) scoperto nella collina di Cagliari nel MDCCCLXVIII. Atti della Reale Accademia Lincei, Memorie della Classe di Scienze Fisiche, Matematiche e Naturali, serie 4 6: 507–533.
• Capellini, G. 1897. A proposito di Tomistoma lyceensis. Rivista Italiana di Paleontologia 3: 18–20.
• Cleuren, J. and De Vree, F. 2000. Feeding in crocodiles. In: K. Schwenk (ed.), Feeding, Form, Function, and Evolution in Tetrapod Vertebrates, 337–358. Academic Press, New York.
• Costa, M., Galobart, A., Roger, E., and Schleich, H. 1995. Primer hallazgo de resto fosiles de cocodrilo en el Parque National de Ordesa y Monte Perdido (Pirineo Central, Huesca).XI Jornadas de Paleontologia, Huesca, 63–64.
• Costa, O.G. 1848. Paleontologia del regno di Napoli. Parte I. Stable Tip. 203 pp. Tramater, Napoli.
• Cuvier, B. 1807. Sur les différentes espèces de crocodiles vivants et sur leurs caractères distinctifs.Annales du Muséum d’Histoire Naturelle 10: 8–66.
• Daudin, F.M. 1801–1803. Histoire naturelle, generale et particuliere des reptiles. Vol. 1. 384 pp. F. Dufart, Paris.
• Del Vecchio, C. 1921. Su alcuni denti di Tomistoma (Crocodilia) dell’Oligocene di Visone presso Acqui. Atti della Società Italiana di Scienze Naturali 60: 419–431.
• Delfino, M. (in press). Late Neogene crocodylian faunas from Libya and the Mediterranean area. In: Geology of East Libya, Proceedings, 3rd Symposium Geology of East Libya, Sedimentary Basins of Libya, 21–23 November 2004. 5 pp. Bingazi, Libya.
• Delfino, M., Pacini, M., Varola, A., and Rook, L. 2003. The crocodiles of the “Pietra Leccese” (Miocene of southern Italy). Abstracts of 1st Meeting European Association of Vertebrate Palaeontology, 18. Basel.
• Delfino, M., Piras, P., and Smith, T. 2005. Anatomy and phylogeny of the gavialoid crocodylian Eosuchus lerichei from the Paleocene of Europe. Acta Palaeontologica Polonica 50: 565–580.
• Dollo, L. 1883. Première note sur les crocodiliens de Bernissart. Bulletin de Musée Royal d’Histoire Naturelles de Belgique 2: 309–340.
• Domning, D.P. and Clark, J.M. 1993. Jamaican Tertiary marine Vertebrata. In: R.M. Wright and E. Robinson (eds.), Biostratigraphy of Jamaica. Geological Society of America Memoir 182: 413–415.
• Efimov, M.B. 1982. New fossil crocodiles from the USSR [in Russian]. Paleontologičeskij žurnal 1982 (2): 146–150.
• Efimov, M.B. 1988. The fossil crocodiles and champsosaurids of Mongolia and USSR [in Russian]. Trudy, Sovmestnaâ Sovetcko−Mongol’skaâ Paleontologičeskaâ Èkspediciâ 36: 1–108.
• Efimov, M.B. 1993. The Eocene crocodiles of the GUS—a history of development. Kaupia 3: 23–25.
• Endo, H., Aoki, R., Taru, H., Kimura, J., Sasaki, M., Yamamoto, M., Arishima K., and Hayashi Y. 2002. Comparative functional morphology of the masticatory apparatus in the long−snouted Crocodiles. Anatomia, Histologia, Embryologia 31: 206–213.
• Erickson, B.R and Sawyer, G.T. 1996. The estuarine crocodile Gavialosuchus carolinensis n. sp. (Crocodylia: Eusuchia) from the Late Oligocene of South Carolina, North America. Science Museum of Minnesota Monographs (Paleontology) 3: 1–47.
• Estes, R. and Báez, A. 1985. Herpetofaunas of North and South America during the Late Cretaceous and Cenozoic: evidence for interchange? In: F.G. Stehli and S.D. Webb (eds.), The Great American Biotic Interchange, 139–197. Plenum Press, New York.
• Fabiani, R. 1915. Il Paleogene del Veneto. Memorie del’Istituto di Geologia della Regia Università di Padova 3: 1–336.
• Falconer, H. and Cautley, P.T. 1840. Notice on the remains of a fossil monkey from the Tertiary strata of the Sewalik Hills in the North of Hindoostan. Transactions of Geological Society of London 2: 499–504.
• Fitzinger, L.J. 1826. Neue Classification der Reptilien nach ihren natürlichen Verwandtschaften. Nebst einer Verwandtschaftstafel und einem Verzeichnisse der Reptiliensammlung des K.K. zoologischen Museum’s zu Wien. vi + 66 pp. Vienna.
• Fourtau, R. 1920. Un nouveau genre de gavialidés. Suppl. II. In: Contribution à l’étude des vertébrés miocènes de l’Égypte, 116–121. Egypt Survey Department, Cairo.
• Gasparini, Z. 1996. Biogeographic evolution of the South American crocodilians. Münchner Geowissenshaftliche Abhandlungen 30: 159–84.
• Gatesy, J., Baker, R.H., and Hayashi, C. 2004. Inconsistencies in arguments for the supertree approach: supermatrices versus supertrees of Crocodylia. Systematic Biology 53: 342–355.
• Geoffroy Saint−Hilaire, E. 1807. Description de deux crocodiles qui existent dans le Nil, compares au crocodile de Saint−Domingue. Annales du Muséum d’Histoire naturelle 10: 67–86.
• Gheerbrant, E., Sudre, J., Iarochene, M., and Moumni, A. 2001. First ascertained African “condylarth” mammals (primitive ungulates: cf. Bulbulodentata and cf. Phenacodonta) from the earliest Ypresian of the Ouled Abdoun Basin, Marocco. Journal of Vertebrate Paleontology 21: 107–118.
• Gilmore, C.W. 1911. A new fossil alligator from the Hell Creek beds of Montana. Proceedings of the United States National Museum 41: 297–302.
• Gmelin, J.F. 1789. Regnum animale. In: Caroli a Linne Systema Naturae per regna tri naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Volume 1 (3), 1033–1516. G.E. Beer, Leipzig.
• Head, J.J. 2001. Systematics and body size of the gigantic, enigmatic crocodyloid Ramphosuchus crassidens, and the faunal history of Siwalik group (Miocene) crocodilians. Journal of Vertebrate Paleontology 21(Supplement to No. 3): 59A.
• Hulke, J.W. 1871. Note on some reptilian fossils from Gozo. Quarterly Journal of Geological Society of London 27: 29–33.
• Huxley, T.H. 1875. On Stagonolepis robertsoni, and on the evolution of the Crocodilia. Quarterly Journal of the Geological Society of London 31: 423–438.
• International Commission on Zoological Nomenclature 1999. International Code of Zoological Nomenclature, Fourth Edition, Adopted by the International Union of Biological Sciences. 306 pp. International Trust for Zoological Nomenclature, London.
• Jonet, S. and Wouters, G. 1977. Maroccosuchus zennaroi, crocodilien eusuchien nouveau des phosphates du Maroc. Notes et Mémoires du Service Géologique du Maroc 38: 177–202.
• Jouve, S. 2004. Etude des crocodyliformes fini Crétace−Paléogène du Bassin de Oulad Abdoun (Maroc) et comparaison avec les faunes africaines contemporaines: systématique, phylogénie et paléobiogéographie. Ph.D. thesis. 652 pp. Muséum National d’Histoire Naturelle de Paris, Paris.
• Kälin, J.A. 1955. Zur Stammesgeschichte der Crocodilia. Revue Suisse de Zoologie 62: 347–56.
• Kamei, T. and Matsumoto, E. 1965. In: N. Kobatake, M. Chiji, N. Ikebe, S. Ishida, T. Kamei, K. Nakaseko, and E. Matsumoto (eds.), Discovery of a Crocodile Fossil from the Osaka Group [in Japanese]. Quaternary Research [= Daiyonki Kenkyu] 4 (2): 47–58.
• Katsura, Y. 2004. Paleopathology of Toyotomaphimeia machikanensis (Diapsida, Crocodylia) from the Middle Pleistocene of Central Japan. Historical Biology 16: 93–97.
• Kobatake, N. and Kamei, T. 1966. The first discovery of fossil crocodile from Central Honshu Japan. Proceedings of Japanese Academy 42: 264–269.
• Kobatake, N., Chiji, M., Ikebe, N., Ishida, S, Kamei, T., Nakaseko, K., and Matsumoto, E. 1965. Discovery of a crocodile fossil from the Osaka Group [in Japanese]. Quaternary Research [= Daiyonki Kenkyu] 4 (2): 47–58.
• Kobayashi, Y. and Tomida, Y. 2005. Anatomy of Toyotomaphimeia machikanensis (Crocodylia) from the Pleistocene of Japan and reassessment of its phylogenetic status. Journal of Vertebrate Paleontology 25 (Supplement to No. 3): 78A–79A.
• Kobayashi, Y., Tomida, Y., Kamei, T., and Eguchi, T. 2006. Anatomy of a Japanese tomistomine crocodylian, Toyotamaphimeia machikanensis (Kamei et Matsumoto, 1965) from the Middle Pleistocene of Osaka prefecture: the reassessment of its phylogenetic status within Crocodylia. National Science Museum Monograph 35: 121.
• Kotsakis, T., Argenti, P., Barisone, G., Delfino, M., Palombo, M.R., Pavia, M., and Piras, P. 2005. Il Paleogene: i vertebrati continentali. In: L. Bonfiglio (ed.), Paleontologia dei Vertebrati in Italia. Memorie del Museo Civico di Storia Naturale di Verona, Serie 2, Scienze della Terra 6: 131–139.
• Kotsakis, T., Delfino, M., and Piras, P. 2004. Italian Cenozoic crocodilians: taxa, timing and biogeographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology 210: 67–87.
• Kuhn, O. 1938. Die Crocodilier aus dem Eozän des Geiseltales bei Halle. Nova Acta Leopoldina, Nova Serie 6: 313–329.
• Lambe, L.M. 1907. On a new crocodilian genus and species from Judith River formation of Alberta. Transactions of the Royal Society of Canada 3: 219–244.
• Langston, W. 1965. Fossil crocodilians from Colombia and the Cenozoic history of the Crocodilia in South America. University of California Publications in Geological Sciences 52: 1–152.
• Langston, W. and Gasparini, Z. 1997. Crocodilians, Gryposuchus and the South American gavials.In: R.F. Kay, R.H. Madden, R.L. Cifelli, and J. Flynn (eds.), Vertebrate Paleontology in the Neotropics. The Miocene fauna of La Venta, Colombia, 113–154. Smithsonian Institution Press, Washington.
• Leidy, J. 1852. Description of a new species of crocodile from the Miocene of Virginia. Journal of Academy of Natural Sciences of Philadelphia 2: 135–138.
• Leslie, A.J. and Taplin, L.E. 2001. Recent development in osmoregulation of crocodilians. In: G. Grigg, F. Seebacher, and C.E. Franklin (eds.), Crocodilian Biology and Evolution, 265–279. Surrey Beatty and Sons, Sydney.
• Li, J.−L. 1975. New material on Tomistoma petrolica of Maoming, Kwantung [in Chinese]. Vertebrata Palasiatica 13: 190–191.
• Llinas−Agrasar, E. 2004. Crocodile remains from the Burdigalian (Lower Miocene) of Gebel Zelten (Libya). Geodiversitas 26: 309–321.
• Lydekker, R. 1886. On the occurrence of the crocodilian genus Tomistoma in the Miocene of the Maltese islands. Quarterly Journal of Geological Society of London 42: 20–22.
• Młynarski, M. 1984 Notes on the amphibian and reptilian faunas of the Polish Miocene. Acta Zoologica Cracoviensia 27: 127–148.
• Mook, C.C. 1924. Further notes on the skull characters of Gavialosuchus americana (Sellards). American Museum Novitates 155: 1–2.
• Mook, C.C. 1955. Two new genera of Eocene crocodilians. American Museum Novitates 1727: 1–4.
• Müller, J. 1838. Waarnemingen over de Indische krokodillen en Beschrijving van eene nieuwe soort. Tydschrift voor Natuurlijke Geschiedenis en Physiologie 5: 67–87.
• Müller, L. 1927. Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Ägyptens. V. Tertiäre Wirbeltiere: 1. Beiträge zur Kenntnis der Krokodilier des ägyptischen Tertiärs. Abhandlungen der Bayerischen Akademie der Wissenschaften 31 (2): 1–97.
• Myrick, A.C. 2001. Thecachampsa antiqua (Leidy, 1852) (Crocodylidae, Thoracosaurinae) from fossil marine deposits at Lee Creek Mine, Aurora, North Carolina, USA. Smithsonian Contributions in Paleobiology 90: 219–225.
• Nicolis, E. 1882. Note illustrative alla carta geologica della provincia di Verona. 150 pp. H.I. Münster, Verona.
• Owen, R. 1850. Monograph on the fossil Reptilia of the London Clay and of the Bracklesham and other Tertiary beds. Part II. Crocodilia. Palaeontographical Society Monographs 3: 5–50.
• Owen, R. 1874. Monograph on the fossil Reptilia of the Wealden and Purbeck Formations. Supplement no. VI. (Hylaeochampsa.). Palaeontographical Society Monographs 27: 1–7.
• Pickford, M. 1994. Late Cenozoic crocodiles (Reptilia: Crocodylidae) from the Western Rift, Uganda. In: B. Senut and M. Pickford (eds.), Geology and Palaeobiology of the Albertine Rift Valley, Uganda−Zaire, Vol. II—Palaeobiology, 137–155. CIFEG, Publication Occasionnelle, Orléans.
• Pinna, G. 1989. Il grande libro dei fossili. 383 pp. Rizzoli, Milano.
• Prothero, D. 1994. The Eocene–Oligocene Transition. 291 pp. Columbia University Press, New York.
• Rage, J.−C. 1996. Terrestrial trans−Tethyan dispersals: an overview. In: Expanded Abstracts of First European Workshop on Vertebrate Palaeontology, 1st–4th May 1996, Copenhagen, 4 pp.
• Rauhe, M., Frey, E., Pemberton, D.S., and Rossmann, T. 1999. Fossil crocodilians from the Late Miocene Baynunah Formation of the Emirate of Abu Dhabi, United Arab Emirates: osteology and palaeoecology. In: J. Whybrow and A. Hill (eds.), Fossil Vertebrates of Arabia, 163–185. Yale University Press, New Haven.
• Roccaforte, P., Sorbini, L., and Piccoli, G. 1994. The fossiliferous sites with Tertiary vertebrates in Northeastern Italy. Memorie di Scienze Geologiche 46: 373–400.
• Rossmann, T. 2002. Studies on Cenozoic crocodiles: 10. First evidence of a tomistomid crocodilian (Eusuchia: Tomistomidae) from the Middle Eocene (Geiseltalian, MP 11) of Grube Messel, Germany. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2002: 129–146.
• Rossmann, T., Berg, D.E., and Salisbury, S.W. 1999. Studies on Cenozoic crocodiles: 3. Gavialosuchus cf. gaudensis (Eusuchia: Tomistomidae) from the Lower Miocene of south Germany. Neues Jahrbuch für Geologie und Paläontologie, Montatshefte 1999: 321–330.
• Sahni, A. and Mishra, V.P. 1975. Lower Tertiary vertebrates from western India. Monographs of the Palaeontological Society of India 3: 1–48.
• Sánchez−Villagra, M.R., Burnham, R.J., Campbell, D.C., Feldmann, R.M., Gaffney, E.S., Kay, R.F., Lozsán, R., Purdy, R., and Thewissen, J.G.M. 2000. A new near−shore marine fauna and flora from the early Neogene of northwestern Venezuela. Journal of Paleontology 74: 957–968.
• Sánchez−Villagra, M.R., Gasparini, Z., Lozsan, R., Moody, J.M. Jr., and Uhen, M.D. 2001. New discoveries of vertebrates from a near−shore marine fauna from the Early Miocene of northwestern Venezuela. Paläontologische Zeitschrift 75: 227–232.
• Scortegagna, F.O. 1836. Sulla formazione geologica della Collina detta la Favorita presso Lonigo. 25 pp. Poligrafica G. Antonelli, Verona.
• Scortegagna, F.O. 1838. Sopra il teschio di un coccodrillo fossile rinvenuto nel Monticello di Lonigo. Atti dell’Ateneo Veneto 2: 139–149.
• Sellards, E.H. 1915. A new gavial from the Late Tertiary of Florida. American Journal of Science 40: 135–138.
• Sellards, E.H. 1916. A new tortoise and a supplementary note on the gavial, Tomistoma americana. American Journal of Science 42: 235–240.
• Shikama, T. 1972. Fossil Crocodilia from Tsochin, south−western Taiwan. Yokohama National University, Science Report, Section II 19: 125–132.
• Souza Filho, J.P. 1991. Charactosuchus sansaoi, uma nova espécie de Crocodilidae (Crocodylia) do Neógeno do Estado o Acre, Brasil. Actas do XII Congreso Brasileiro de Paleontologia, 36. São Paulo.
• Souza Filho, J.P. and Bocquentin, J. 1989. Brasilosuchus mendensi n.g., n.sp., um novo representante da familia Gavialidae do Neógeno do Estado do Acre, Brasil. Anais do XI Congreso Brasiliero de Paleontologia 1: 457–463. Curitiba.
• Souza Filho, J.P., Bocquentin, J., and Silva, E. 1993. Novas interpretaçoes do genero Brasilosuchus (Crocodylia) do Neógeno do Estado do Acre, Brasil. Resumos do XIII Congreso Brasileiro de Paleontologia e do I Simposio de Paleontologia Cone Sul, 130. São Leopoldo.
• Steel, R. 1973. Handbuch der Paläoherpetologie, Vol. 16: Crocodylia. 116 pp. Fischer−Verlag, Stuttgart.
• Sun, A., Li, J., Ye, X., Dong, Z., and Hou, L. 1992. The Chinese Fossil Reptiles and Their Kins. 260 pp. Science Press, Beijing.
• Swinton, W.E. 1937. The crocodile of Maransart (Dollosuchus dixoni [Owen]). Mémoires du Musée d’Histoire Naturelle de Belgique 80: 1–44.
• Swofford, D.L. 2002. PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods). Version 4. Sinauer Associates, Sunderland.
• Taruno, H. 1999. A fossil crocodile from Nagareki Town, Kishiwada City. In: Kishiwada City Board of Education (ed.), Excavation Report on a Fossil Crocodile from Nagareki Town, Kishiwada City, 1–36. Natural History Museum of Kishiwada City, Kishiwada, Osaka.
• Tchernov, E. 1986. Evolution of the crocodiles in East and North Africa. Cahier de Paléontologie. Travaux de Paléontologie est−africaine. 65 pp. Editions CNRS, Paris.
• Toula, F. and Kail, J.A. 1885. Über einen Krokodil−Schädel aus den Tertiärablagerungen von Eggenburg in Niederösterreich: eine paläontologische Studie. Denkschriften der Keiserlichen Akademie der Wissenschaften von Wien, Mathematisch−naturwissenschaftliche Klasse 50: 299–355.
• Turner, A. and Antón, M. 2004. Evolving Eden. 269 pp. Columbia University Press, New York.
• Uzielli, G. 1886. Sopra un cranio di coccodrillo trovato nel Modenese. Bollettino della Società Geologica Italiana 5: 355–364.
• Vasse, D. 1993. Systématique de crocodiles du Crétacé supérieur et du Paléogène d’Europe. Aspects paléobiogéographiques et paléoécologiques. Unpublished Ph.D. thesis. 224 pp. Université Paris VI, Paris.
• Vianna, A. and Moraes, A. 1945. Sur un crâne de crocodile fossile decouvert dans le Miocène de Lisbonne. Boletim de la Sociedade Geológica de Portugal 4: 161–170.
• Vijayasarathi, G. and Sabale, A.B. 1985. Miocene crocodilian (Tomistomidae) fossil from southwestern Kutch, Gujarat.Current Science 54: 1071–1073.
• Webb, S.D. and Tessman, N. 1967. Vertebrate evidence of a low sea level in the middle Pliocene. Science 156: 379.
• West, R.M., Hutchinson, J.H., and Munthe, J. 1991. Miocene vertebrates from the Siwalik Group, Western Nepal. Journal of Vertebrate Paleontology 11: 108–129.
• Wood, S.V. 1846. On the discovery of an alligator and of several new Mammalia in the Hordwell Cliff; with observations upon the geological phenomena of that locality. London Geological Journal 1: 117–122.
• Yeh, H.−K. 1958. A new crocodile from Maoming, Kwangtung [in Chinese]. Vertebrata Palasiatica 2: 237–242.
• Zigno, A. de 1880. Sopra un cranio di coccodrillo scoperto nel terreno eoceno del Veronese. Atti della Reale Accademia Lincei, Memorie della Classe di Scienze Fisiche, Matematiche e Naturali, Serie 3 5: 65–72.