A new ankylosaurid from the Late Cretaceous Two Medicine Formation of Montana, USA

• Autorzy: Penkalski P.
• Języki publikacji: EN

Abstrakty

EN

Oohkotokia horneri gen. et sp. nov. is described based on a specimen in the collections of the Museum of the Rockies, Montana, USA. Oohkotokia exhibits a unique combination of characters not seen in other late Campanian North American ankylosaurids: prominent, horn-like, trihedral squamosal bosses, a small, undistinguished median nasal plate on the dorsal surface of the rostrum, a relatively small occipital condyle, a smooth, finely pitted osteoderm external texture, and triangular lateral osteoderms. Other specimens from the Two Medicine Formation are referable to Oohkotokia. O. horneri, Euoplocephalus tutus, Dyoplosaurus acutosquameus, and Scolosaurus cutleri separate stratigraphically.

Słowa kluczowe

EN

ankylosaur; Dinosauria; Ornithischia; Ankylosauridae; systematics; paleontology; Campanian; Late Cretaceous; Cretaceous; Two Medicine Formation; Montana; USA

• Wydawca: -
• Czasopismo: Acta Palaeontologica Polonica
• Rocznik: 2014
• Tom: 59
• Numer: 3
• Opis fizyczny: p.617-634,fig.,ref.

Twórcy

autor

Penkalski P.

• The Geology Museum, 1215 W. Dayton St., Madison, WI 53706, USA

Bibliografia

• Arbour, V.M. 2010. A Cretaceous armoury: multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA. Journal of Vertebrate Paleontology 30 (Supplement 3): 55A.
• Arbour, V.M. and Snively, E. 2009. Finite element analyses of ankylosaurid dinosaur tail club impacts. The Anatomical Record 292: 1412–1426.
• Arbour, V.M. and Currie, P.J. 2012. Analyzing taphonomic deformation of ankylosaur skulls using retrodeformation and finite element analysis. PloS One 7 (6): e39323.
• Arbour, V.M. and Currie, P.J. 2013. Euoplocephalus tutus and the diversity of ankylosaurid dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USA. PLoS ONE 8 (5): e62421.
• Arbour, V.M., Burns, M.E., and Sissons, R. 2009. A redescription of the ankylosaurid dinosaur Dyoplosaurus acutosquameus Parks, 1924 (Ornithischia: Ankylosauria) and a revision of the genus. Journal of Vertebrate Paleontology 29: 1117–1135.
• Arbour, V.M., Burns, M.E., and Currie, P.J. 2011. A review of pelvic shield morphology in ankylosaurs (Dinosauria: Ornithischia). Journal of Paleontology 85: 298–302.
• Bakker, R.T. 1988. Review of the Late Cretaceous nodosauroid Dinosauria. Hunteria 1 (3): 1–23.
• Béland, P. and Russell, D.A. 1978. Paleoecology of Dinosaur Provincial Park (Cretaceous), Alberta, interpreted from the distribution of articulated vertebrate remains. Canadian Journal of Earth Sciences 15: 1012–1024.
• Bell, P.R. 2012. Standardized terminology and potential taxonomic utility for hadrosaurid skin impressions: a case study for Saurolophus from Canada and Mongolia. PLoS ONE 7 (2): e31295.
• Blows, W.T. 2001. Dermal armor of the polacanthine dinosaurs. In: K. Carpenter (ed.), The Armored Dinosaurs, 363–385. Indiana University Press, Bloomington.
• Brown, B. 1908. The Ankylosauridae, a new family of armoured dinosaurs from the Upper Cretaceous. Bulletin of the American Museum of Natural History 24: 187–201.
• Burnham, D.A., Derstler, K.L., Currie, P.J., Bakker, R.T., Zhou, Z., and Ostrom, J.H. 2000. Remarkable new birdlike dinosaur (Theropoda: Maniraptora) from the Upper Cretaceous of Montana. University of Kansas Paleontological Contributions 13: 1–14.
• Burns, M.E. 2008. Taxonomic utility of ankylosaur (Dinosauria, Ornithischia) osteoderms: Glyptodontopelta mimus Ford, 2000: a test case. Journal of Vertebrate Paleontology 28: 1102–1109.
• Burns, M.E. and Sullivan, R.M. 2011. A new ankylosaurid from the Upper Cretaceous Kirtland Formation, San Juan Basin, with comments on the diversity of ankylosaurids in New Mexico. New Mexico Museum of Natural History and Science Bulletin 53: 169–178.
• Burns, M.E., Currie, P.J., Sissons, R.L., and Arbour, V.M. 2011. Juvenile specimens of Pinacosaurus grangeri Gilmore, 1933 (Ornithischia: Ankylosauria) from the Late Cretaceous of China, with comments on the specific taxonomy of Pinacosaurus. Cretaceous Research 32: 174–186.
• Carpenter, K. 1990. Ankylosaur systematics: Example using Panoplosaurus and Edmontonia (Ankylosauridae: Nodosauridae). In: K. Carpenter and P.J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives, 281–298. Cambridge University Press, Cambridge.
• Carpenter, K. 2001. Phylogenetic analysis of the Ankylosauria. In: K. Carpenter (ed.), The Armored Dinosaurs, 455–483. Indiana University Press, Bloomington.
• Carpenter, K. 2004. Redescription of Ankylosaurus magniventris Brown 1908 (Ankylosauridae) from the Upper Cretaceous of the Western Interior of North America. Canadian Journal of Earth Sciences 41: 961–986.
• Carpenter, K., Bartlett, J., Bird, J., and Barrick, R. 2008. Ankylosaurs from the Price River Quarries, Cedar Mountain Formation (Lower Cretaceous), East-Central Utah. Journal of Vertebrate Paleontology 28: 1089–1101.
• Carpenter, K., Hayashi, S., Kobayashi, Y., Maryańska, T., Barsbold, R., Sato, K., and Obata, I. 2011. Saichania chulsanensis (Ornithischia, Ankylosauridae) from the Upper Cretaceous of Mongolia. Palaeontographica, Abteilung A 294: 1–61.
• Carpenter, K., Kirkland, J.I., Birge, D., and Bird, J. 2001. Disarticulated skull of a new primitive ankylosaurid from the Lower Cretaceous of Utah. In: K. Carpenter (ed.), The Armored Dinosaurs, 211–238. Indiana University Press, Bloomington.
• Chinnery, B.J. and Horner, J.R. 2007. A new neoceratopsian dinosaur linking North American and Asian taxa. Journal of Vertebrate Paleontology 27: 625–641.
• Coombs, W.P., Jr. 1971. The Ankylosauria. 487 pp. Unpublished Ph.D. dissertation, Columbia University, New York.
• Coombs, W.P., Jr. 1978. The families of the ornithischian dinosaur order Ankylosauria. Palaeontology 21: 143–170.
• Coombs, W. P., Jr. 1986. A Juvenile ankylosaur referable to the genus Euoplocephalus (reptilia, ornithischia). Journal of Vertebrate Paleontology 6: 162–173.
• Coombs, W.P., Jr. 1990. Teeth and taxonomy in ankylosaurs. In: K. Carpenter and P.J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives, 269–279. Cambridge University Press, Cambridge.
• Coombs, W.P., Jr. 1995. Ankylosaurian tail clubs of middle Campanian to early Maastrichtian age from western North America, with description of a tiny club from Alberta and discussion of tail orientation and tail club function. Canadian Journal of Earth Sciences 32: 902–912.
• Coombs, W.P., Jr. and Demere, T.A. 1996. A Late Cretaceous nodosaurid ankylosaur (Dinosauria: Ornithischia) from marine sediments of coastal California. Journal of Paleontology 70: 311–326.
• Currie, P.J. 2003. Cranial anatomy of tyrannosaurid dinosaurs from the Late Cretaceous of Alberta, Canada. Acta Palaeontologica Polonica 48: 191–226.
• Dawson, G.M. 1885. Report on the region in the vicinity of the Bow and Belly Rivers, North West Territory. Canada Geological Survey Report of Progress 1882-83-84: 1–169c.
• Dodson, P. 1997. Paleoecology. In: P.J. Currie and K. Padian (eds.), Encyclopedia of Dinosaurs, 515–519. Academic Press, San Diego.
• Dong, Z.M. 2002. A new armored dinosaur (Ankylosauria) from Beipiao Basin, Liaoning Province, northeastern China. Vertebrata PalAsiatica 40: 276–285.
• Eberth, D.A. 1997. Judith River wedge. In: P.J. Currie and K. Padian (eds.), Encyclopedia of Dinosaurs, 379–385. Academic Press, San Diego.
• Eberth, D.A. 2005. The geology. In: P.J. Currie and E.B. Koppelhus (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed, 54–82. Indiana University Press, Bloomington.
• Eberth, D.A. and Braman, D.R. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053–1086.
• Ford, T.L. and Kirkland, J.I. 2001. Carlsbad ankylosaur (Ornithischia, Ankylosauria): an ankylosaurid and not a nodosaurid. In: K. Carpenter (ed.), The Armored Dinosaurs, 239–260. Indiana University Press, Bloomington.
• Gilmore, C.W. 1917. Brachyceratops, a ceratopsian dinosaur from the Two Medicine Formation of Montana, with notes on associated fossil reptiles. U.S. Geological Survey Professional Paper 103: 1–45.
• Gilmore, C.W. 1930. On dinosaurian reptiles from the Two Medicine Formation of Montana. Proceedings of the United States National Museum 77 (16): 1–39.
• Gilmore, C.W. 1939. Ceratopsian dinosaurs from the Two Medicine Formation, Upper Cretaceous of Montana. Proceedings of the U.S. National Museum 87: 1–18.
• Godefroit, P., Pereda-Suberbiola, X., Li, H., and Dong, Z.M. 1999. A new species of the ankylosaurid dinosaur Pinacosaurus from the Late Cretaceous of Inner Mongolia (P.R. China). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre 69 (supplement): 17–36.
• Hamblin, A.P. 1994. The Dinosaur Park Formation of the Upper Cretaceous Judith River (Belly River) Group, subsurface of southern Alberta. Geological Survey of Canada Open File 2831.
• Hayashi, S., Carpenter, K., Scheyer, T.M., Watabe, M., and Suzuki, D. 2010. Function and evolution of ankylosaur dermal armor. Acta Palaeontologica Polonica 55: 213–228.
• Hieronymus, T.L, Witmer, L.M., Tanke, D.H., and Currie, P.J. 2009. The facial integument of centrosaurine ceratopsids: Morphological and histological correlates of novel skin structures. The Anatomical Record 292: 1370–1396.
• Hill, R.V. and Norell, M.A. 2008. Pinacosaurus grangeri comes of age: ontogenetic changes in the ankylosaurian skull. Journal of Vertebrate Paleontology 28: 91A.
• Hill, R.V., Witmer, L.M., and Norell, M.A. 2003. A new specimen of Pinacosaurus grangeri (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia: Ontogeny and phylogeny of Ankylosaurs. American Museum Novitates 3395: 1–29.
• Horner, J.R. 1982. Evidence of colony nesting and “site fidelity” among ornithischian dinosaurs. Nature 297: 675–676.
• Horner, J.R. 1983. Cranial osteology and morphology of the type specimen of Maiasaura peeblesorum (Ornithischia: Hadrosauridae) with discussion of its phylogenetic position. Journal of Vertebrate Paleontology 3: 29–38.
• Horner, J.R. 1984. Three ecologically distinct vertebrate faunal communities from the Late Cretaceous Two Medicine Formation of Montana, with discussion of evolutionary pressures induced by interior seaway fluctuations. In: J.D. McBane and P. Garrison (eds.), Northwest Montana and Adjacent Canada, Montana Geological Society Field Conference, 299–304. Montana Geological Society, Billings.
• Horner, J.R. 1999a. Dinosaur biozones in the Campanian clastic wedge complex of Montana, and their biological implications. American Association of Petroleum Geologists Bulletin 83: 1184.
• Horner, J.R. 1999b. Egg clutches and embryos of two hadrosaurian dinosaurs. Journal of Vertebrate Paleontology 19: 607–611.
• Horner, J.R. and Makela, R. 1979. Nest of juveniles provides evidence of family structure among dinosaurs. Nature 282: 296–298.
• Horner, J.R. and Weishampel, D.B. 1988. A comparative embryological study of two ornithischian dinosaurs. Nature 332: 256–257.
• Horner, J.R., Varricchio, D.J., and Goodwin, M.B. 1992. Marine transgressions and the evolution of Cretaceous dinosaurs. Nature 358: 59–61.
• Kirkland, J.I. 1998. A polacanthine ankylosaur (Ornithischia: Dinosauria) from the Early Cretaceous (Barreminan) of Eastern Utah. New Mexico Museum of Natural History and Science Bulletin 14: 271–281.
• Lambe, L.M. 1902. New genera and species from the Belly River Series (Mid-Cretaceous). Contributions to Canadian Palaeontology, Geological Survey of Canada 3: 25–81.
• Lambe, L.M. 1910. Note on the parietal crest of Centrosaurus apertus, and a proposed new generic name for Stereocephalus tutus. Ottawa Naturalist 14: 149–151.
• Lü, J., Ji, Q., Gao, Y., and Li, Z. 2007. A new species of the ankylosaurid dinosaur Crichtonsaurus (Ankylosauridae: Ankylosauria) from the Cretaceous of Liaoning Province, China. Acta Geologica Sinica 81: 883–897.
• Maleev, E.A. 1952. A new ankylosaur from the Upper Cretaceous of Mongolia [in Russian]. Doklady Akademii Nauk S.S.S.R. 87: 273–276.
• Mallon, J.C., Evans, D.C., Ryan, M.J., and Anderson, J.S. 2012. Megaherbivorous dinosaur turnover in the Dinosaur Park Formation (upper Campanian) of Alberta, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology 350–352: 124–138.
• Maryańska, T. 1977. Ankylosauridae (Dinosauria) from Mongolia. Palaeontologia Polonica 37: 85–151.
• McDonald, A.T. and Horner, J.R. 2010. New material of “Styracosaurus” ovatus from the Two Medicine Formation of Montana. In: M.J Ryan, B.J. Chinnery-Allgeier, and D.A. Eberth (eds.), New Perspectives on Horned Dinosaurs, 156–168. Indiana University Press, Bloomington.
• Nopcsa, F. 1918. Leipsanosaurus n. gen. ein neuer thyreophore aus der Gosau. Földtani Közlöny 48: 324–328.
• Nopcsa, F. 1928. Paleontological notes on reptiles. Geologica Hungarica, Series Palaeontologica 1: 51–74.
• Osborn, H.F. 1923. Two Lower Cretaceous dinosaurs of Mongolia. American Museum Novitates 95: 1–10.
• Ostrom, J.H. 1970. Stratigraphy and paleontology of the Cloverly Formation (Lower Cretaceous) of the Bighorn Basin Area, Wyoming and Montana. Peabody Museum of Natural History Bulletin 35: 1–234.
• Owen, R. 1842. Report on British fossil reptiles. Part II. Report British Association for the Advancement of Science 1841: 60–204.
• Pang, Q. and Cheng, Z. 1998. A new ankylosaur of late Cretaceous from Tianzhen, Shanxi. Progress in Natural Science 8: 326–334.
• Parks, W.A. 1924. Dyoplosaurus acutosquameus: a new genus and species of armoured dinosaur with notes on a skeleton of Prosaurolophus maximus. University of Toronto Studies (Geological Series) 18: 1–35.
• Parsons, W.L., and Parsons, K.M. 2009. A new ankylosaur (Dinosauria: Ankylosauria) from the Lower Cretaceous Cloverly Formation of central Montana. Canadian Journal of Earth Sciences 46: 721–738.
• Penkalski, P. 1998. A preliminary systematic analysis of Ankylosauridae (Ornithischia: Thyreophora) from the Late Campanian of North America. Journal of Vertebrate Paleontology 18 (Supplement 3): 69A–70A.
• Penkalski, P. 2001. Variation in specimens referred to Euoplocephalus tutus. In: K. Carpenter (ed.), The Armored Dinosaurs, 261–298. Indiana University Press, Bloomington.
• Penkalski, P. and Blows, W.T. 2013. Scolosaurus cutleri (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada. Canadian Journal of Earth Sciences 50: 171–182.
• Rogers, R.R. 1997. Two Medicine Formation. In: P.J. Currie and K. Padian (eds.), Encyclopedia of Dinosaurs, 760–765. Academic Press, San Diego.
• Rogers, R.R., Swisher III, C.C., and Horner, J.R. 1993. 40Ar/39Ar age and correlation of the nonmarine Two Medicine Formation (Upper Cretaceous), northwestern Montana, U.S.A. Canadian Journal of Earth Sciences 30: 1066–1075.
• Ryan, M.J. and Evans, D.C. 2005. Late Cretaceous ceratopsids from the Judith River Group, Alberta, and the recognition of dinosaurian faunal zones within Dinosaur Provincial Park. In: D.R. Braman, F. Therrien, E.B. Koppelhus, and W. Taylor (eds.), Dinosaur Park Symposium—Short Papers, Abstracts, and Program, 92–93. Royal Tyrrell Museum, Drumheller.
• Ryan, M.J., Holmes, R., and Russell, A.P. 2007. A revision of the Late Campanian centrosaurine ceratopsid genus Styracosaurus from the western interior of North America. Journal of Vertebrate Paleontology 27: 944–962.
• Sampson, S.D. 1995. Two new horned dinosaurs from the Upper Cretaceous Two Medicine Formation of Montana; with a phylogenetic analysis of the Centrosaurinae (Ornithischia: Ceratopsidae). Journal of Vertebrate Paleontology 15: 743–760.
• Seeley, H.G. 1888. The classification of the Dinosauria. Report British Association for the Advancement of Science 1887: 698–699.
• Stebinger, E. 1914. Stratigraphy of the Montana group (Upper Cretaceous) in northwestern Montana and its relation to the Belly River beds and Judith River Formation. Journal of the Washington Academy of Sciences 4: 383–384.
• Sternberg, C.M. 1929. A toothless armored dinosaur from the Upper Cretaceous of Alberta. Bulletin of the National Museum of Canada 54: 28–33.
• Sullivan, R.M. 1999. Nodocephalosaurus kirtlandensis, gen. et sp. nov, a new ankylosaurid dinosaur (Ornithischia: Ankylosauria) from the Upper Cretaceous Kirtland Formation (Upper Campanian), San Juan Basin, New Mexico. Journal of Vertebrate Paleontology 19: 126–139.
• Sullivan, R.M. and Fowler, D.W. 2006. New specimens of the rare ankylosaurid dinosaur Nodocephalosaurus kirtlandensis (Ornithischia: Ankylosauridae) from the Upper Cretaceous Kirtland Formation (Dena-zin Member), San Juan Basin, New Mexico. New Mexico Museum of Natural History and Science Bulletin 35: 259–261.
• Sullivan, R., Arbour, V., Burns, M., and Lucas, S. 2013. A new ankylosaurid dinosaur (Ornithischia, Ankylosauria) from the Upper Cretaceous Kirtland Formation, San Juan Basin, New Mexico, USA. Journal of Vertebrate Paleontology 33 (Supplement): 222.
• Swofford, D.L. 2003. PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods). Version 4. Sinauer Associates, Sunderland.
• Tanoue, K. 2005. Postcranial skeleton and ontogeny of Euoplocephalus tutus (Ornithischia, Ankylosauridae). Journal of Vertebrate Paleontology 25 (Supplement 3): 121A.
• Thompson, R.S., Parish, J.C., Maidment, S.C.R., and Barrett, P.M. 2012. Phylogeny of the ankylosaurian dinosaurs (Ornithischia: Thyreophora). Journal of Systematic Palaeontology 10: 301–312.
• Trexler, D. 2001. Two Medicine Formation, Montana: geology and fauna. In: D.H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life, 298–309. Indiana University Press, Bloomington.
• Tumanova, T.A. 1987. The armored dinosaurs of Mongolia [in Russian]. Trudy Sovmestnaâ Sovecko-Mongolskaâ Paleontologičeskaâ Ekspedicâ 32: 1–80.
• Tumanova, T.A. 2000. Armoured dinosaurs from the Cretaceous of Mongolia. In: M.J. Benton, E.N. Kurochkin, M.A. Shishkin, and D.M. Unwin (eds.), The Age of Dinosaurs in Russia and Mongolia, 517–532. Cambridge University Press, Cambridge.
• Varricchio, D.J., Horner, J.R., and Jackson, F.D. 2002. Embryos and eggs for the Cretaceous theropod dinosaur Troodon formosus. Journal of Vertebrate Paleontology 22: 564–576.
• Vickaryous, M.T. and Russell, A.P. 2003. A redescription of the skull of Euoplocephalus tutus (Archosauria: Ornithischia): a foundation for comparative and systematic studies of ankylosaurian dinosaurs. Zoological Journal of the Linnean Society 137: 157–186.
• Vickaryous, M.T., Russell, A.P., and Currie, P.J. 2001. Cranial ornamentation of ankylosaurs: reappraisal of developmental hypotheses. In: K. Carpenter (ed.), The Armored Dinosaurs, 318–340. Indiana University Press, Bloomington.
• Vickaryous, M.K., Maryańska, T., and Weishampel, D.B. 2004. Ankylosauria. In: D.B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria (2nd edition), 363–392. University of California Press, Berkeley.
• Weishampel, D.B., Barrett, P.M., Coria, R.A., Le Loueff, J., Xu, X., Zhao, X., Sahni, A., Gomani, E.M.P., and Noto, C. 2004. Dinosaur distribution. In: D.B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria (2nd edition), 517–606. University of California Press, Berkeley.
• Xu, L., Lü, J.C., Zhang, X.L., Jia, S.H., Hu, W.Y., Zhang, J.M., Wu, Y.H., and Ji, Q. 2007. New nodosaurid ankylosaur from the Cretaceous of Ruyang, Henan Province. Acta Geologica Sinica 81: 433–438.

Identyfikatory

DOI

2012.0125