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2015 | 73 |

Tytuł artykułu

Changes in horizontal structure of natural beech forests on an altitudinal gradient in the Sudetes

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Języki publikacji



The article describes horizontal structure of the tree layer, natural regeneration, snags and crown projections of natural beech stands on three permanent research plots in the wide altitudinal range in the Krkonoše Mts (Czech Republic). The spatial structure was classified from 1980 to 2010 and subsequently the prediction of spontaneous development with an outlook for 30 years (to 2040) was done by growth simulator. Hopkins-Skellam index, Pielou-Mountford index, Clark-Evans index and Ripley’s K-function were calculated. Further, the vertical structure and total diversity index was evaluated. The horizontal structure of individuals in the tree layer had not changed significantly during the monitored years. Tree spatial pattern of the lowest altitude lying herb-rich beech forest was mostly regular to random, in acidophilous mountain beech forest predominantly random and in fragments of beech groups around the timberline aggregated. Juvenile growth on all investigated plots was distributed aggregated and snags randomly. The horizontal structure of crown projection centroids had always higher values toward the regularity than tree layer and was random to regular. The result of principal component analysis also confirmed that spatial pattern was dependent on the altitude, but also on the number of trees.







Opis fizyczny



  • Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 129, 165 21 Prague, Czech Republic
  • Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 129, 165 21 Prague, Czech Republic
  • Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 129, 165 21 Prague, Czech Republic
  • Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamycka 129, 165 21 Prague, Czech Republic
  • National Forest Centre, T.G.Masaryka 22, 460 01 Zvolen, Slovak Republic


  • Barrera M.D., Frangi J.L., Richter L.L., Perdomo M.H., Pinedo L.B. 2000. Structural and functional changes in Nothofagus pumilio forests along an altitudinal gradient in Tierra del Fuego, Argentina. Journal of Vegetation Science 11: 179–188.
  • Christensen M., Hahn K., Mountford E.P., Odor P., Standovar T., Rozenbergar D., Diaci J., Wijdeven S., Meyer P., Winter S., Vrska T. 2005. Dead wood in European beech (Fagus sylvatica) forest reserves. Forest Ecology and Management 210: 267–282.
  • Clark P.J., Evans F.C. 1954. Distance to nearest neighbor as a measure of spatial relationships in populations. Ecology 35: 445–453.
  • Closset-Kopp D., Schnitzler A., Aran D. 2006. Dynamics in natural mixed-beech forest of the Upper Vosges. Biodiversity and Conservation 15: 1063–1093.
  • Commarmot B., Bachofen H., Bundziak Y., Bürgi A., Ramp B., Shparyk Y., Sukhariuk D., Viter R., Zingg A. 2005. Structures of virgin and managed beech forests in Uholka (Ukraine) and Sihlwald (Switzerland): a comparative study. Forest Snow and Landscape Research 79: 45–56.
  • Crookston N.L., Stage A.R. 1999. Percent canopy cover and stand structure statistics from the Forest Vegetation Simulator. US Department of Agriculture, Forest Service, Rocky Mountain Research Station.
  • Doležal J., Šrůtek M. 2002. Altitudinal changes in composition and structure of mountain-temperate vegetation: a case study from the Western Carpathians. Plant Ecology 158: 201–221.
  • Ellenberg H. 1996. Vegetation Mitteleuropas mit den Alpen. 5.Aufl. Ulmer, Stuttgart.
  • Fabrika M. 2005. Simulátor biodynamiky lesa SIBYLA, koncepcia, konštrukcia a programové riešenie. Technická univerzita vo Zvolene.
  • Ferris-Kaan R., Peace A.J., Humphrey J.W. 1998. Assessing structural diversity in managed forests. In: Assessment of Biodiversity for Improved Forest Planning. European Forest Institute Proceedings 18. Bachmann P. (ed.). Kluwer Academic Publishers, p. 331–342.
  • Füldner K. 1995. Strukturbeschreibung in Mischbeständen. Forstarchiv 66: 235–606.
  • Geyer C.J. 1999. Likelihood Inference for Spatial Point Processes. Chapter 3 In: Geometry: Likelihood and Computation. Barndorff-Nielsen O.E., Kendall W.S., Van Lieshout M.N.M. (eds.). Stochastic Chapman and Hall/CRC, Monographs on Statistics and Applied Probability 80: 79–140.
  • Girardin C.A.J., Farfan-Rios W., Garcia K., Feeley K.J., Jørgensen P.M., Murakami A.A., Perez L.C., Seidel R., Paniagua N., Fuentes Claros A.F., Maldonado C., Silman M., Salinas N., Reynel C., Neill D.A., Serrano M., Caballero C.J., de los Angeles La Torre Cuadros M., Macia M.J., Killeen T.J., Malhi Y. 2014. Spatial patterns of above-ground structure, biomass and composition in a network of six Andean elevation transects. Plant Ecolology and Diversity 7: 161–171.
  • Holdridge L.R. 1967. Life zone ecology. Tropical Science Center. San Jose, Costa Rica.
  • Holeksa J., Saniga M., Szwagrzyk J., Dziedzic T., Ferenc S., Wodka M. 2007. Altitudinal variability of stand structure and regeneration in the subalpine spruce forests of the Poľana biosphere reserve, Central Slovakia. European Journal of Forest Research 126: 303–313.
  • Hopkins B., Skellam J.G. 1954. A new method for determining the type of distribution of plant individuals. Annals of Botany 18: 213–227.
  • Jaehne S.C., Dohrenbusch A. 1997. Ein Verfahren zur Beurteilung der Bestandesdiversität. Forstwissenschaftliches Centralblatt 116: 333–345.
  • Jeník J. 1998. Biodiversity of the Hercynian mountains of Central Europe. Pirineos 151/152: 83–99.
  • Kooijman A.M., Emmer I.M., Fanta J., Sevink J. 2000. Natural regeneration potential of the degraded Krkonoše forests. Land Degradation and Development 11: 459–473.
  • Korpeľ Š. 1995. Die Urwälder der Westkarpaten. Stuttgart, Jena, New York, Gustav Fischer Verlag.
  • Le Tacon F. 1981. Caractérisation edaphique. In: Le hêtre. Teissier du Cros E., Le Tacon F., Nepveu G., Pardé J., Perrin R., Timbal J. (eds.). INRA, Paris, pp. 77–84.
  • McElhinny C., Gibbons P., Brack C., Bauhus J. 2005. Forest and woodland stand structural complexity: its definition and measurement. Forest Ecology and Management 218: 1–24.
  • Ministry of Agriculture of the Czech Republic 2013. Report on the state of forests and forestry in the Czech Republic by 2013.
  • Motta R., Morales M., Nola P. 2006. Human land-use, forest dynamics and tree growth at the treeline in the Western Italian Alps. Annals of Forest Science 63: 739–747.
  • Mountford E.P. 2002. Fallen dead wood levels in the near-natural beech forest at La Tillaie reserve, Fontainebleau, France. Forestry 75: 203–208.
  • Mountford M.D. 1961. On E. C. Pielou’s index of non-randomness. Journal of Ecology 49: 271–275.
  • Nagel T.A., Svoboda M., Diaci, J. 2006. Regeneration patterns after intermediate wind disturbance in an old-growth Fagus–Abies forest in southeastern Slovenia. Forest Ecology and Management 226: 268–278.
  • Neuhäuslová Z., Blažková D., Grulich V., Husová M., Chytrý M., Jeník J., Jirásek J., Kolbek J., Kropáč Z., Ložek V., Moravec J., Prach K., Rybníček K., Rybníčková E., Sádlo J. 1998. Mapa potenciální přirozené vegetace České republiky. Textová část. (Map of the Potential Natural Vegetation of the Czech Republic. Explanatory text.). Academia, Praha.
  • Neumann M., Starlinger F. 2001. The significance of different indices for stand structure and diversity in forests. Forest Ecology and Management 145: 91–106.
  • Otto H.J. 1994. Waldökologie. Stuttgart, Ulmer.
  • Pittner J., Saniga M. 2008. A change in structural diversity and regeneration processes of the spruce virgin forest in Nefcerka NNR (TANAP) in relation to altitude. Journal of Forest Science 54: 545–553.
  • Pretzsch H. 1992. Konzeption und Konstruktion von Wuchsmodellen für Rein- und Mischbestände. Forstliche Forschungsberichte, München.
  • Pretzsch H. 2001. Modellierung des Waldwachstums. Parey Buchverlag, Berlin.
  • Pretzsch H. 2009. Forest Dynamics, Growth and Yield. Springer.
  • Rademacher C., Neuert C., Grundmann V., Wissel C., Grimm V. 2001. Was charakterisiert Buchenurwälder? Untersuchungen der Altersstruktur des Kronendachs und der raümlichen Verteilung der Baumriesen in einem Modellwald mit Hilfe des Simulationsmodells BEFORE. Forstwissenschaftliches Centralblatt 120: 288–302.
  • Ripley B. D. 1977. Modelling Spatial Patterns. Journal of the Royal Statistical Society. Series B 39: 2: 172–212.
  • Rozas V. 2006. Structural heterogeneity and tree spatial patterns in old-growth deciduous lowland forest in Cantabria, northern Spain. Plant Ecology 185: 57–72.
  • Rugani T., Diaci J., Hladnik D. 2013. Gap Dynamics and Structure of Two Old-Growth Beech Forest Remnants in Slovenia. PloS one 8: e52641.
  • Schröter M., Härdtle W., von Oheimb G. 2012. Crown plasticity and neighborhood interactions of European beech (Fagus sylvatica L.) in an old-growth forest. European Journal of Forest Research 131: 787–798.
  • Šmelko Š. 2007. Dendrometria. 2nd release. Zvolen. TU Zvolen.
  • Špulák O., Souček J. 2010. The Sibyla model and development of beech forests affected by air pollution. Central European Journal of Biology 5: 371–383.
  • Štefančík I. 2013. Effect of delayed tending on development of beech (Fagus sylvatica L.) pole stage stand. Folia Oecologica 40: 272–281.
  • ter Braak C.J.F., Šmilauer P. 2002. CANOCO Reference Manual and CanoDraw for Windows User’s Guide: Software for Canonical Community Ordination (version 4.5). Ithaca, NY, USA ( Microcomputer Power.
  • Vacek S., Nosková I., Bílek L., Vacek Z., Schwarz O. 2010a. Regeneration of forest stands on permanent research plots in the Krkonoše Mts. Journal of Forest Science 56: 11: 541–554.
  • Vacek S., Vacek Z., Schwarz O., Raj A., Bilek L., Noskova I., Balcar Z., Zahradnik D., Balas M., Bednarik J., Mikesa M., Minx T., Matejka K. 2010b. Struktura a vývoj lesních porostů na výzkumných plochách v národních parcích Krkonoš. Kostelec nad Černými lesy: Lesnická práce.
  • Vacek S., Hejcman M. 2012. Natural layering, foliation, fertility and plant species composition of a Fagus sylvatica stand above the alpine timberline in the Giant (Krkonoše) Mts., Czech Republic. European Journal of Forest Research 131: 799–810.
  • Vera F.W.M. 2000. Grazing ecology and forest history. CABI Publishing, Oxon.
  • von Oheimb G., Westphal C., Härdtle, W. 2007. Diversity and spatio-temporal dynamics of dead wood in a temperate near-natural beech forest (Fagus sylvatica). European Journal of Forest Research 126: 359–370.
  • von Oheimb G., Westphal C., Tempel H., Härdtle W. 2005. Structural pattern of a near-natural beech forest (Fagus sylvatica) (Serrahn, north-east Germany). Forest Ecology and Management 212: 253–263.
  • Weber J. 1999. Beschreibung der Diversität von Bestandesstrukturen mit Hilfe von Teilkollektiven. Tagung des Deutschen Vereins Forstlicher Forschungsanstalten. Sektion Forstliche Biometrie und der AG Okologie der Internationalen Biometrischen Gesellschaft in Freiburg, pp. 25–37.
  • Wijdeven S.M.J. 2004. Stand dynamics in Fontainebleau. Dynamics in beech forest structure and composition over 17 years in La Tillaie forest reserve, Fontainebleua, France. Green World Research, Alterra.
  • Wolf A. 2005. Fifty year record of change in tree spatial patterns within a mixed deciduous forest. Forest Ecology and Management 215: 212–223.
  • Zeibig A., Diaci J., Wagner S. 2005. Gap disturbance patterns of a Fagus sylvatica virgin forest remnant in the mountain vegetation belt of Slovenia. Forest Snow and Landscape Research 79: 69–80.

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