Lochkovian conodonts from Podolia, Ukraine, and their stratigraphic significance

• Autorzy: Drygant D. , Szaniawski H.
• Języki publikacji: EN

Abstrakty

EN

In the Podolian Dniester Basin (southwestern Ukraine) the Lower Devonian marine deposits are represented by about 530 m thick continuous sequence of interlaminated carbonate and schale outcrops at several localities. Conodonts occur in most of the carbonate layers of the whole Lochkovian but are not abundant and their ramiform elements are mostly broken or lacking. Therefore, only the pectiniform, Pa elements of twenty five stratigraphically important conodont species occurring in the region are discussed and two new species, Caudicriodus schoenlaubi and Pandorinellina? parva are proposed. The hypothetical phyletic relationships within the main representatives of the icriodontid and spathognathodontid genera, Caudicriodus, Zieglerodina, and Pandorinellina? are traced. Comparison of the previously published and newly obtained data revealed discrepancies in the hitherto used interpretation of some of the conodont taxa and their stratigraphic ranges. Contrary to the earlier reports, Caudicriodus postwoschmidti does not occur in the lower Lochkovian but only in the middle part of the Chortkiv Formation, high above the Monograptus uniformis Zone. Based on new material and verification of the previous determinations, a modified scheme of the Lochkovian conodont zonation in Podolia is proposed. Conodont zones: Caudicriodus hesperius, C. transiens, C. postwoschmidti, C. serus, and ?Caudicriodus steinachensis are distinguished. The zones are correlated with conodont zonations in other regions—Barrandian, Cantabrian Mountains, Pyrenees, and Nevada. Biostratigraphy of the Siluro−Devonian transition and Lochkovian is integrated with the carbon isotope stratigraphy.

Słowa kluczowe

EN

Lochkovian; conodont; Podolia; Ukraine; stratigraphy; Conodonta; evolution; Devonian; paleontology

• Wydawca: -
• Czasopismo: Acta Palaeontologica Polonica
• Rocznik: 2012
• Tom: 57
• Numer: 4
• Opis fizyczny: p.833-861,fig.,ref.

Twórcy

autor

Drygant D.

• State Museum of Natural History, National Academy of Sciences of Ukraine, Teatralna 18, Lviv 79008, Ukraine

autor

Szaniawski H.

Bibliografia

• Al−Rawi, D. 1977. Biostratigraphische Gliederung der Tentaculiten–Schichten des Frankenwaldes mit Conodonten und Tentaculiten (Unter− und Mittel−Devon; Bayern, Deutschland). Senckenbergiana lethaea 58: 25–79.
• Baliński, A. 2010. First colour−patterned strophomenidae brachiopod from the earliest Devonian of Podolia, Ukraine. Acta Palaeontologica Polonica 55: 695–700.
• Baliński, A. 2012. The brachiopod succession through the Silurian–Devonian boundary beds at Dnistrovo, Podolia, Ukraine. Acta Palaeontologica Polonica 57: 897–924.
• Barrick, J.E. and Klapper, G. 1992. Late Silurian–Early Devonian Conodonts from the Hunton Group (Upper Henryhouse, Haragan, and Bois d’Arc Formations), South−Central Oklahoma. Oklahoma Geological Survey Bulletin 145: 19–65.
• Barrick, J.E., Meyer, B.D., and Ruppel, S.C. 2005. The Silurian–Devonian boundary and the Klonk event in the Frame Formation, subsurface West Texas. Bulletins of American Paleontology 369: 105–122.
• Broadhead, T.W. and McComb, R. 1983. Paedomorphosis in the conodont family Icriodontidae and the evolution of Icriodus. Fossils and Strata 15: 149–154.
• Buggisch, W. and Mann, U. 2004. Carbon isotope stratigraphy of Lochkovian to Eifelian limestones from the Devonian of central and southern Europe. International Journal of Earth Sciences (Geologische Rundschau) 93: 521–541.
• Bultynck, P. 1976. Le Silurien Superieur et le Devonien Inferieur de la Sierra de Guadarrama (Espagne Centrale). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen 49 (5): 1–73.
• Bultynck, P. 2003. Devonian Icriodontidae: biostratigraphy, classification and remarks on paleoecology and dispersal. Revista Española de Micropaleontología 35: 295–314.
• Carls, P. 1969. Die Conodonten des tieferen Unter−Devons der Guadarrama (Mittel−Spanien) und die Stellung der Grenzbereiches Lochkovium/Pragium nach der rheinischen Gliederung. Senckenbergiana lethaea 50: 303–355.
• Carls, P. 1975. Zusätzliche Conodonten−Funde aus dem tieferen Unter−Devon Keltiberiens (Spanien). Senckenbergiana lethaea 56: 399–429.
• Carls, P. and Gandl, J. 1969. Stratigraphie und Conodonten des Unter-Devons der Östlichen Iberischen Ketten (NE−Spanien). Neues Jahbuch für Geologie und Paläontologie, Abhandlungen 132: 155–218.
• Carls, P., Slavík, L., and Valenzuela−Ríos, J.I. 2007. Revisions of conodont biostratigraphy across the Silurian–Devonian boundary.Czech Geological Survey, Bulletin of Geosciences 82: 145–164.
• Chatterton, B.D.E. and Perry, D.G. 1977. Lochkovian trilobites and conodonts from northwestern Canada. Journal of Paleontology 51: 772–796.
• Chlupáč, I. and Hladil, J. 2000. The global stratotype section and point of the Silurian–Devonian boundary. Courier Forschungsinstitut Senckenberg 225: 1–7.
• Colquhoun, G.P. 1995. Early Devonian conodont faunas from the Capertee High, NE Lachlan Fold Belt, southeastern Australia.Courier Forschungsinstitut Senckenberg 182: 347–369.
• Corradini, C. and Corriga, M.G. 2012. A Přídolí–Lochkovian conodont zonation in Sardinia and the Carnic Alps: implication for a global zonation scheme. Bulletin of Geosciences 87: 635–650.
• Drygant, D.M. 1967. Some data on the Conodont Zones and the age of the Silurian–Devonian passage beds in Podolia [in Russian]. Paleontologičeskij Sbornik 4 (2): 56–59.
• Drygant, D.M. 1968a. Conodont zones of the Ludlovian and Gedinnian equivalents in Podolia [in Russian]. Referaty dokladov k III Meždunarodnomu simpozûmu po granice silura i devona i stratigrafii nižnego i srednego devona, 70–72. All−Union Geological Scientific Research Institute (VSEGEI), Leningrad.
• Drygant, D.M. 1968b. Some conodont species from the Podolian Silurian [in Russian]. Paleontologičeskij Sbornik 5 (1): 46–52.
• Drygant, D.M. 1971 Conodont zones of the Ludlovian and Gedinnian equivalents in Podolia [in Russian]. In: D.V. Nalivkin (ed.), Granica silura i devona i biostratigrafiâ silura. Trudy III Meždunarodnogo simposûma, Leningrad, 1968 1: 85–89.
• Drygant, D.M. 1974. Simple Silurian and Lower Devonian conodonts from the Volyn−Podolia [in Russian]. Paleontologičeskij Sbornik 10 (2): 64–70.
• Drygant, D.M. 1984. Korrelaciâ i konodonty silurijskih−nižniedevonskih otloženij Volyno−Podolii. 192 pp. Naukova Dumka, Kiyev.
• Drygant, D.M. 1994. About the Conodont zonation in Lower Devonian [in Ukrainian]. Paleontolohičny Zbirnyk 30: 38–42.
• Drygant, D.M. 2000. Lower and Middle Paleozoic of the Volyn’−Podillja margin of the East European Platform and Carpathian Foredeep [in Ukrainian]. Proceedings of the State Natural History Museum, Lviv 15: 24–130.
• Drygant, D.M. 2010. Devonian Conodonts from South−West Margin of the East European Platform (Volyn'−Podolian Ukraine) [in Ukrainian]. 156 pp. Academperiodyka, Kyiv.
• Drygant, D.M. and Szaniawski, H. 2008. Conodont stratigraphy of the Lower Devonian in Podolia, Ukraine. In: P. Königshof (ed.), Field Workshop 2008 of the IGCP 499−UNESCO “Devonian Land−Sea Interaction: Evolution of Ecosystems and Climate (DEVEC)”, 33. Libyan Petroleum Institute and Forschungsinstitut und Naturmuseum Senckenberg, Franfut am Mein.
• Drygant, D. and Szaniawski, H. 2009. Conodonts of the Silurian–Devonian boundary beds in Podolia, Ukraine. In: M.G. Corriga and S. Piras (eds.), Time and Life in the Silurian: a multidisciplinary approach. Rendiconti della Società Paleontologica Italiana 3: 281–282.
• Farrell, J.R. 2003. Late Přidoli, Lochkovian, and early Pragian conodonts from the Gap area between Larras Lee and Eurimbla, central western NSW, Australia. Courier Forschungsinstitut Senckenberg 245: 107–181.
• Filipiak, P., Zatoń, M., Szaniawski, H., and Wrona, R. 2012. Palynology and microfacies of Lower Devonian mixed carbonate−siliciclastic deposits in Podolia, Ukraine. Acta Palaeontologica Polonica 57: 863–877.
• García−López, S., Jahnke, H., and Sanz−López, J. 2002. Uppermost Pridoli to Upper Emsian stratigraphy of the Alto Carrión Unit, Palentine Domain (Northwest Spain). In: S. García−López and F. Bastida (eds), Palaeozoic Conodonts from Northern Spain. Instituto Geológico y Minero de España, serie Cuadernos del Museo Geominero 1: 229–257.
• Jeppsson, L. 1989. Latest Silurian Conodonts from Klonk, Czechoslovakia. Geologica et Palaeontologica 23: 21–37.
• Johnson, D.B. and Klapper, G. 1981. New Early Devonian conodont species of central Nevada. Journal of Paleontology 55: 1237–1250.
• Klapper, G. 1969. Lower Devonian conodont sequence, Royal Creek, Yukon Territory, and Devon Island, Canada. Journal of Paleontology 43: 1–27.
• Klapper, G. 1977. With contributions by D.B. Johnson. Lower and Middle Devonian conodont sequence in central Nevada. In: M.A. Murphy, W.B.N. Berry, and C.A. Sandberg, (eds.), Western North America: Devonian. Univesity of California, Riverside Campus Museum Contribution 4: 33–54.
• Klapper, G. and Johnson, J.G. 1980. Endemism and dispersal of Devonian conodonts. Journal of Paleontology 54: 400–455.
• Klapper, G. and Murphy, M.A. 1975. Silurian–Lower Devonian Conodont Sequence in the Roberts Mountains Formation of central Nevada. University of California Publications in Geological Sciences 111: 1–62.
• Klapper, G. and Ziegler, W. 1979. Devonian conodont biostratigraphy. In: M.R. House, C.T. Scrutton, and M.G. Basset (eds.), The Devonian System. Special Papers in Palaeontology 23: 199–224.
• Kozłowski, R. 1929. Les Brachiopodes Gothlandiens de la Podolie Polonaise. Palaeontologia Polonica 1: 1–254.
• Lane, H.R. 1968. Symmetry in conodont element−pairs. Journal of Paleontology 42: 1258–1263.
• Lane, H.R. and Ormiston, A.R. 1979. Siluro−Devonian biostratigraphy of the Salmontrout River area, east−central Alaska. Geologica et Palaeontologica 13: 39–96.
• Małkowski, K., Racki, G., Drygant, D., and Szaniawski, H. 2009. Carbon isotope stratigraphy across the Silurian–Devonian transition in Podolia, Ukraine: evidense for a global biogeochemical perturbation. Geological Magazine 146: 674–689.
• Mashkova, T.V. [Maškova, T.V.] 1968a. Some Conodonts of the Borshchiv and Chortkiv Horizons of the Podolia [in Russian]. In: B.S. Sokolov (ed.), Stratigrafiâ nižnego paleozoâ Centralnoj Evropy: Meždunarodnyj geologičeskij kongress. Doklady soveckih geologov, 145–148. Nauka, Moskva.
• Mashkova, T.V. [Maškova, T.V.] 1968b. The Conodonts of the genus Icriodus Branson et Mehl, 1938 from the Borshchiv and Chortkiv Horizons of Podolia (in Russian). Doklady AN SSSR 182: 941–944.
• Mashkova, T.V. [Maškova, T.V.] 1970. On the range of the woschmidti Zone in Podolia (in Russian). Doklady AN SSSR 190: 654–657.
• Mashkova, T.V. [Maškova, T.V.] 1971. Zonal conodont assemblages from boundary beds of the Silurian and Devonian of Podolia [in Russian]. In: D.V. Nalivkin (ed.), Granica silura i devona i biostratigrafiâ silura. Trudy III Meždunarodnogo simpozûma. Leningrad, 1968 1: 157–164.
• Mashkova, T.V. 1972. Ozarkodina steinhornensis (Ziegler) apparatus, its conodonts and biozone. Geologica et Palaeontologica SB 1: 81–91.
• Mashkova, T.V. 1979. Conodont zones of the Lower Devonian in the U.S.S.R. Geologica et Palaeontologica 13: 97–102.
• Mawson, R. 1986. Early Devonian (Lochkovian) conodont faunas from Windellama, New South Wales.Geologica et Palaeontologica 20: 39–71.
• Mawson, R. and Talent, J. 1994. Age of an Early Devonian carbonate fan and isolated limestone clasts and megaclasts, east−central Victoria. Proceedings of the Royal Society of Victoria 106: 31–70.
• Mawson, R., Talent, J.A., Molloy, P., and Simpson, A.J. 2003. Siluro−Devonian (Pridoli–Lochkovian and early Emsian) conodonts from the Nowshera area, Pakistan: implications for the mid−Palaeozoic stratigraphy of the Peshawar Basin. Courier Forschungsinstitut Senckenberg 245: 83–105.
• Murphy, M.A. and Cebecioglu, M.K. 1984. The Icriodus steinachensis and I. claudiae lineages (Devonian Conodonts). Journal of Paleontology 58: 1399–1411.
• Murphy, M.A. and Matti, J.G. 1982. Lower Devonian Conodonts (hesperiuskindlei Zones), Zentral Nevada. University of California Publications in Geological Sciences 123: 1–83.
• Murphy, M.A. and Valenzuela−Ríos, J.I. 1999. Lanea new genus of Early Devonian conodonts. Bollettino della Società Paleontologica Italiana 37: 321–334.
• Murphy, M.A., Valenzuela−Ríos, J.I., and Carls, P. 2004. On classification of Pridoli (Silurian)–Lochkovian (Devonian) Spathognathodontidae (conodonts). University of California, Riverside Campus Museum Contribution 6: 1–25.
• Nikiforova, O.I. 1977. Podolia. The Silurian–Devonian Boundary. IUGS, Series A 5: 52–64.
• Nikiforova, O.I. and Priedtiechiensky, N.N. 1968. A guide to the geological excursion on Silurian and Lower Devonian deposits of Podolia (Middle Dniestr River). 58 pp. The Ministry of Geology of the USSR−All−Union Geological Scientific Research Institute (VSEGEI), Leningrad.
• Nikiforova, O.I., Priedtiechiensky, N.N. [Priedtiečensky, N.N.], Abushik, A.F. [Abušik, A.F.], Ignatovich, M.M. [Ignatovič, M.M], Modzalevskaya, T.L. [Modzalevskaâ, T.L.], Berger, A.Ya. [Berger, A.Â], Novoselova, L.S., Burkov, Yu.K. [Burkov, Û.K.] 1972. Opornyj razriez silura i nižniego devona Podolii. 262 pp. Nauka, Leningrad.
• Olivieri, R. and Serpagli, E. 1990. Latest Silurian–early Devonian conodonts from the Mason Porcus Section near Fluminimaggiore, South−western Sardinia. Bollettino della Società Paleontologica Italiana 29: 59–76.
• Olempska, E. 2012. Exceptional soft−tissue preservation in boring ctenostome bryozoans and associated “fungal” borings from the Early Devonioan of Podolia, Ukraine. Acta Palaeontologica Polonica 57: 925–940.
• Olempska, E., Horne D.J., and Szaniawski, H. 2011. First record of preserved soft parts in Paleozoic podocopid (Metacopina) ostracod, Cytherellina submagna: phylogenetic implications. Proceedings of the Royal Society Biological Sciences 279: 564–570.
• Philip, G.M. 1965. Lower Devonian conodonts from the Tyers Area, Gippsland, Victoria.Proceedings of the Royal Society of Victoria 79: 95–117.
• Racki, G., Baliński, A., Wrona, R., Małkowski, K., Drygant, D., and Szaniawski, H. 2012. Faunal dynamics across the Silurian–Devonian positive isotope excursion (δ¹³C, δ¹⁸O) in Podolia, Ukraine: Comparative analysis of the Ireviken and Klonk events. Acta Palaeontologica Polonica 57: 795–832.
• Savage, N.M. 1982. Lower Devonian (Lochkovian) conodonts from Lulu Island, Southeastern Alaska. Journal of Paleontology 56: 983–988.
• Schönlaub, H.P. 1980a. Silurian and Devonian conodont localities of the Barrandian. Field Trip E. Second European Conodont Symposium (ECOS II). Abhandlungen der Geologischen Bundesanstalt 35: 147–180.
• Schönlaub, H.P. 1980b. Carnic Alps. Field Trip A. Second European Conodont Symposium (ECOS II). Abhandlungen der Geologischen Bundesanstalt 35: 5–57.
• Schönlaub, H.P. 1985. Conodonts. In: I. Chlupáč, P. Lukeš, F. Paris, and H.P. Schönlaub (eds.), The Lochkovian–Pragian Boundary in the Lower Devonian of the Barrandian Area (Czechoslovakia). Jahrbuch der Geologischen Bundesanstalt 128: 9–41.
• Serpagli, E. 1983. The conodont apparatus ofIcriodus woschmidti woschmidti Ziegler. Fossils and Strata 15: 155–161.
• Simpson, A. 1998. Apparatus structure of the latest Silurian to Early Devonian conodont Icriodus woschmidti hesperius Klapper et Murphy, and some comments on phylogeny. In: H. Szaniawski (ed.), Proceedings of the Sixth European Conodont Symposium (ECOS VI). Palaeontologia Polonica 58: 153–169.
• Simpson, A. 2003. A new subspecies of the conodont genus Ozarkodina and its correlative value.Courier Forschungsinstitut Senckenberg 245: 75–81.
• Slavík, L. 2004. A new zonation of the Pragian Stage (Lower Devonian) in the stratotype area (Barrandian, central Bohemia). Newsletters on Stratigraphy 40: 39–71.
• Slavík, L. and Hladil, J. 2004. Lochkovian/Pragian GSSP revisited: evidence abaut conodont taxa and their stratigraphic distribution. Newsletters on Stratigraphy 40: 137–153.
• Slavík, L., Carls, P., Hladil, J., and Koptíková, L. (in press). Subdivision of the Lochkovian Stage based on conodont faunas from the stratotype area (Prague Synform, Czech Republic). Geological Journal.
• Slavík, L., Valenzuela−Ríos, J.I., Hladil, J., and Carls, P. 2007. Early Pragian conodont−based correlations between the Barrandian area and the Spanish Central Pyrenees. Geological Journal 42: 499–512.
• Suttner, T.J. 2007. Conodont stratigraphy, Facies−Related Distribution Patterns and Stable Isotopes (Carbon and Oxygen) of the Uppermost Silurian to Lower Seewarte Section (Carnic Alps, Carinthia, Austria). Abhandlungen der Geologischen Bundesanstalt 59: 1–111.
• Sweet, W.C. and Schönlaub, H.P. 1975. Conodonts of the Genus Oulodus Branson and Mehl, 1933. Geologica et Palaeontologica 9: 4159.
• Szajnocha, W. 1889. O stratygrafii pokładów sylurskich galicyjskiego Podola. Sprawozdania Komisyi Fizyograficznej Akademii Umiejętności 23: 185–200.
• Uyeno, T.T. 1990. Biostratigraphy and conodont faunas of Upper Ordovician through Middle Devonian rocks, eastern Arctic Archipelago. Geological Survey of Canada, Bulletin 401: 1–211.
• Uyeno, T.T. 1991. Pre−Famennian Devonian conodont biostratigraphy of selected intervals in the eastern Canadian Cordillera. Geological Survey of Canada, Bulletin 417: 129–161.
• Valenzuela−Ríos, J.I. 1994. Conodontos del Lochkoviense y Praguiense (Devónico Inferior) del Pirineo Central Español. Memorias del Museo Paleontülogicode la universidad de Zaragoza 5: 1–178.
• Valenzuela−Ríos, J.I. and Murphy, M.A. 1997. A new zonation of middle Lochkovian (Lower Devonian) conodonts and evolution of Flajsella n. gen. (Conodonta). In: G. Klapper, M.A. Murphy, and J.A. Talent (eds.), Paleozoic Sequence Stratigraphy, Biostratigraphy and Biogeography, Studies in Honor of J.Granville (“Jess”) Johnson. Boulder, Colorado. Geological Society of America Special Paper 321: 131–144.
• Viira, V. 1999. Late Silurian conodont biostratigraphy in the northern East Baltic. Bollettino della Società Paleontologica Italiana 37: 299–310.
• Voichyshyn, V.K. [Voyčyšyn, V.K.] 2001. Distribution of fossil rimains of Agnatha and accompanying vertebrate groups in Lower Devonian deposits of Podillia [in Ukrainian]. Naukovi zapysky Deržavnoho pryrodoznavčoho muzeyu) 16: 47–58.
• Voichyshyn, V.K. 2011. The Early Devonian Armoured Agnathans of Podolia, Ukraine. Palaeontologia Polonica 66: 1–211.
• Voichyshyn, V.K. and Szaniawski, H. 2012. Acanthodian jaw bones from Lower Devonian marine deposits of Podolia, Ukraine. Acta Palaeontologica Polonica 57: 879–896.
• Walliser, O.H. 1960. Scolecodonts, conodonts, and vertebrates. In: A.J. Boucot, A. Martinsson, R. Thorsteinsson, O.H. Walliser, H.B. Whittington, and E. Yochelson (eds.), A Late Silurian fauna from the Sutherland River formation, Devon Islands, Canadian Arctic Archipelago. Geological Survey of Canada, Bulletin 65: 1–51.
• Walliser, O.H. 1964. Conodonten des Silurs. Abhandlungen des Hessischen Landesamtes für Bodenforschung 41: 1–106.
• Weddige, K. 1996. Devon–Korrelationstabelle. Senckenbergiana Lethaea 76: 267–286.
• Ziegler, W. 1960. Conodonten aus dem Rheinischen Unterdevon (Gedinnium) des Remscheider Sattels (Rheinisches Schiefergebirge). Paläontologische Zeitschrift 34: 169–201.
• Ziegler, W. (ed.) 1975. Catalogue of Conodonts. Vol. 2. 404 pp. E. Schwiezerbart'sche Verlagsbuchhandlung, Stuttgart.